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Book Title
Versuche und Beobachtungen über die Bastarderzeugung im Pflanzenreich
By
Publication Details
Stuttgart, [s.n.], 1849
DOI
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Cambridge University Library
Sponsor
JISC & NEH
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Darwin Estate and Cambridge University Library

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In copyright


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top-margin annotation I think began Sept 15 /54/
show subjects concepts

lines 3—2 annotation in dark brown ink 1849


lines 14—10 score
line 11 underline "Uebereinkunft"
lines 13—1 annotation great agreemt in animals & Plants in Bastardsng
show subjects concepts


line 21 score "XVII" in pale pencil


line 15 underline "Vereinigung"
line 15 annotation joining


lines 12—14 score in pale pencil
lines 12—17 annotation cause of failure chiefly in female. organs.


lines 1—2 score

line 5 underline "Zutritts"
line 5 annotation access
line 7 underline "Versehen"
line 7 annotation oversight
line 8 underline "anderswoher"
line 8 annotation elsewhere
line 8 underline "irgend"
line 8 annotation some

lines 17—15 annotation cancelled Herbert. p 371 )

line 22 underline "schliesst"
line 22 annotation prevents/

lines 11—8 annotation all injurious influences more injurious to hybridising.


lines 11—12 score
line 13 underline "häufig ... Blume"
line 14 underline "bloss"
line 12 annotation naked
top-marginline 2 annotation N.B. As damp & rain so injurious to fructification , it makes it odder that flowers are not ‹comm› regularly impregnated in closed state, for thy can be impregnated haufig in this condition. In cases of Campanula which are impregnated in bud, are these foreigners? & wd. thy open in own country.
show subjects subjects

line 15 underline "gewissermassen"
line 15 annotation so much or more
line 15 underline "bastardartig"
line 15 annotation of the nature of bastards
show subjects concepts

lines 12—8 score
line 6 underline "entartete:
line 6 annotation degenerated
line 4 underline "flexibel"
bottom-margin annotation flexible or plastic
lines 3—2 score


lines 11—12 score
lines 9—12 annotation some effect of variability on hybrids /

lines 14—17 score
lines 14—19 annotation no great difference in hybridisation of wild & cultivated . /
lines 14—16 annotation in brown ink (Q)

line 19 underline "Erzeugung"
line 19 annotation production

[continues overleaf] lines 9—5 score
lines 9—5 annotation Disputes Herberts' case of fertile Hybrids
line 2 underline "vorhandene"
line 2 annotation existing
lines 3—1 score
line 3bottom-margin annotation thinks has mistaken th fertility of some hybrids, with th results of a first impregnation.


[continuation] 1 underline "Es ... aber"
line 1 annotation on the other hand
line 3 underline "unmittelbaren"
line 3 annotation immediate
lines 2—6 annotation some hybrid fruits are richer in seed, than the fruit produced by first union. /

line 6 underline "entschiedene"
line 6 annotation decided
line 9 underline "begreifen"
line 9 annotation comprehend
show subjects concepts

line 14 underline "die ... Typen"
line 14 annotation (?)
show subjects concepts

line 10 underline "Uebereinstimmung"
line 10 annotation accordance
line 6 underline "Gährung"
line 6 annotation fermenting


line 7 underline in pale pencil "umständlicher"
line 7 annotation circumstantial


line 5 underline "Wesen" @5 annotation existence
line 10 underline "vollkommenen"
line 10 annotation perfect
line 5 underline "zeugen"
line 5 annotation witness.
line 3 underline "Verkümmerung"
line 3 annotation arresting
line 3 underline "Staubgefässe"
line 3 annotation anthers


line 3 underline "Verletzungen"
line 3 annotation wounds


line 16 underline "verschwinden"
line 16 annotation disappear

lines 13—6 annotation condition of pollen on stigma changes sooner or later according to relationship


line 1 underline "Täuschung"
bottom-margin annotation illusion


line 9 underline "kehren"
line 9 annotation return


line 5 underline "Zeitraum"
line 5 annotation period


lines 17—21 annotation fruit falls off, from an imperfect impregnation


line 6 underline in pale pencil "Vertheilung"
line 6 annotation distribution


top-marginline 15 annotation It is not Hybrids — but Hybrid-fructification .    Most important distinction which I have ovrlooked.

lines 18—21 double score
lines 17—21 annotation Hybrids never produce full amont of seeds

line 21 underline "ungeachtet"
line 21 annotation ‹howeve› «notwithstan»


lines 7—10 annotation Hybridisation requires all outward circumstances favourable.
show subjects subjects


line 7 underline "fremden"
line 7 annotation Never the least effect .


line 13 at "Pollenarten" annotation no mixed effect.—

line 5 underline "rustica"
line 5 annotation female
line 5 annotation prefers paniculata

line 4 at "N. paniculata" annotation female
line 3 at "rustica" annotation prefers Langsdorf
line 3 underline "Langsdorfii"


line 11 underline "erfordert"
line 11 annotation requires
line 10 underline "Beziehung"
line 10 annotation reference


lines 5—6 multiple score


lines 1—7 score
line 3 underline Ansätze ... Zähne"
top-marginline 7 annotation case of variety with characters like other species    (Q)


line 10 underline "dreifach ... gemischten"
lines 8—10 annotation 3 — sorts in same capsules


lines 18—22 annotation cancelled Herbert believes in tincture


lines 7—13 score
top-marginline 16 annotation This is what might have been expected
     mere physical difficulty ??

lines 12—11 score
line 14 underline "Befruchtung ... Pollen"
lines 20—10 annotation This slowness is important as it is character in parents & not in hybrid.

lines 8—6 score
lines 7—5 annotation injures the other ovules:


line 7 underline "fülich"
line 7 annotation convenient


line 10 underline "einem ... Pollen"
line 12 underline "rustico-paniculata"
line 12 underline "Langsdorfii"
lines 7—13 annotation In Hybrids father or mother's pollen makes own powerless. so does quite foreign pollen sometimes

line 15 underline "zumal"
line 15 annotation specially
line 16 underline "exotischen"
line 20 underline "Lobelia"
line 20 annotation Example
lines 15—22 annotation In some pure, specially exotics, own pollen will not ‹fe› impregnate, whilst that of other individual, or even other species, yet own pollen good.—
show subjects subjects

line 9 underline "B ... Herbert"
line 9 underline "Zephyranthes"
lines 10—6 annotation p. 355t01 so Herbert with Zephyranthes but not good example; see XX next Page .
t01 - `p. 355' in dark pencil

line 5 at "Hippeastrum" annotation ⧟ Amaryllidaet01 p. 371— in this case it is Hybrid with hybrid. —
t01 - `Amaryllidae' in dark pencil

line 5 underline "Bosse"
lines 4—3underline "Passiflora"


line 3 underline "veträgt"
line 3 annotation accord

lines 5—9 score
line 6 annotation XX
top-margin annotation XX This like Herberts Zephyranthes case; Pas. racemosa can be fructified by pollen of coerulea, but stigm of coerulea cannot be fructified by pollen of racemosa or by its own. — we may say female organ of coerulea injured .

lines 10—12 score
lines 10—12 annotation takes the view given above XX
show subjects subjects

line 13 underline "Einrichtung"
line 13 annotation disposition
lines 14—15 underline "Parietaria judaica"
lines 14—16 annotation vertically crossed Has not heard of Conrad Sp
line 14 at "Parietaria" annotation ⧟ (not in Sprengel)
line 19 underline "zipfeln"
line 19 annotation tips
lines 21—23 score
lines 21—23 annotation compares with Snails

line 13 underline "Bedingung" annotation condition


lines 9—23 annotation / Thinks th fruit of Hybrids is not due to pollen-influence, but to that power of forming fruit, which th most sterile hybrids without any pollen do produce
     Repeated p. 106
show subjects subjects


lines 1—3 score
lines 1—3 annotation mother not more powerful than pollen

lines 13—15 annotation Hybridising generally no effect on seeds.

lines 14—12 annotation apple half sour half sweet.—

line 8 underline "Birnbäumchen" /@line 8 annotation pear
line 7 at "verschiedener" annotation various
line 7 underline "Obstarten"
line 7 annotation fruit
line 7 underline "liess"
line 4 underline " Räthselhaftes"
line 4 annotation enigmatical
lines 8—3 annotation castrated pear-blossoms in orchard bore much fruit, showing crossing
line 1 score
bottom-margin annotation thinks experiment not careful enough.


line 10 underline "geneigt"
line 10 annotation inclined

bottom-margin annotation Disbelieves (perhaps vry truly) all these cases of direct effect of pollen on th mothers' fruit.—


line 11 underline "Pfropfen"
line 11 annotation graft

lines 19—18 annotation in brown ink Q
line 18 underline "Pelargonium zonale"
line 18 annotation sport in
lines 14—13 underline "Levcojen ... Cheiri"
line 14 annotation sports in
line 9 underline "Achillaea Millefolium"
line 9 annotation sports in wild
line 9 underline "weissen"
lines 9—8 underline "stark ... Blumendolden"
line 8 annotation umbels


line 2 underline "fleissigen"
line 2 annotation diligent
line 5 underline "zuverlässig"
line 5 annotation authentic
line 9 underline "Andeutung"
line 9 annotation notice


bottom-margin annotation see p. 499 . & p 135


lines 9—6 double score
top-margin6 annotation cancelled This most important, ‹either› «if» crossing varieties ‹does nothing to seeds, or› «has had anything to do with diverse coloured» seeds , then they are crossed naturally by Bees . —
show subjects subjects

line 11 annotation X
show subjects subjects

lines 14—13 annotation all a mistake
lines 12—9 score
lines 7—6 score
line 6 underline "reinen"
line 9bottom-margin annotation cancelled The self-impregnated flowers gave same result as the castrated & cross impregnated & these showed colours altered

bottom-margin annotation cancelled Has tried Wiegman experiments with quite different results, ie seeds never affected see next several Pages


line 1 underline "verwendet"
line 1 annotation employed
lines 1—2 underline "lauter ... Gattung"
lines 1—2 annotation most constant vars.

lines 19—25 score
lines 19—25 annotation here seeds were coloured

line 8 score
line 8 annotation cancelled were these mongrels


[continues overleaf] 2—3 underline "grünlich-gelben"
line 5 underline "gelben"
line 10 underline "mehr grünlich"
line 16 underline "grünen"
line 3 underline "gelben"


[continuation] 1 underline "schmützig-grüne"/ @line 1 annotation color

lines 20—24 score
line 22 underline "entschieden"
line 22 annotation decided
from End Note annotation Ask Author
     p. 84    Were any of «th» mongrel Peas reared ?


lines 10—11 double score
line 10 underline "kommen"
line 10 annotation agree
line 11 underline "Wesentlichen"
line 11 annotation essentiality
lines 10—11 annotation (& Berkeleys)

line 13 underline "Piso-Vicia"
line 13 annotation see account    p. 14
line 15 underline "geringer Fruchtbarkeit"

lines 19—14 annotation plants from Wiegmanns Pisum-Vicia he ‹rather› thinks it a
line 13 underline "Varietäts-Bastard"
line 13bottom-margin annotation «rather» than hybrid, because, ⧟
line 8 at "Mit dieser Pflanze" annotation ie offsrping of Wiegmans Piso-Vicia
line 5bottom-margin annotation flower impregnated with common pea , produced same result as self.impregn, & pollen of Vicia had no effect .


line 12 underline "sondern ... war"
lines 13—10 annotation Conclusion mere variety, & says nothing about mongrel .

lines 7—4 score
line 7 underline "multiflorus"
line 6 underline "nanus"
line 6 underline "americanus"
lines 7—4 annotation cd not make any of them cross.
from End Note annotation (86 dirct actn of pollen)

line 6 at "nanus" annotation ⧟ Loudon makes these distinct species, besides vulgaris—
show subjects concepts


line 1 underline "Misslingen"
line 1 annotation failure

lines 6—8 score
lines 6—8 annotation Leguminosae opposed to Hybridisation
show subjects subjects

line 9 score
lines 13—15 double score
lines 8—16 annotation If then mongrelising takes place easily, yet cannot at all believe in Wiegman hybrids.

line 18 at "annua" annotation Cruciferae
line 20 underline "getrauen"
line 20 annotation dare
lines 18—21 annotation doubts about seeds in Matthiola what to say
line 22 underline "gegenseitigen"
line 22 annotation mutual
line 25 at "anders" annotation otherwise
line 26 underline "herrührt"
line 26 annotation comes from

lines 9—8 score
lines 5—1 score
line 3 underline "einer ... die"
bottom-margin annotation Mays not affected


top-margin annotation also Berkeley did not artificially cross.—}—


line 9 annotation XX
line 12 underline "bestritten"
line 12 annotation contest?
line 15 at "sonst" annotation besides
line 21 at "berechtigt" annotation authorise
top-margin annotation XX It seems he does «fully» admit that cross fructification does in Pisum affect seeds, & as Wiegmann did not artificially impregnate, shows that Peas, when ‹found› «not castrated» are crossed «naturally.» — Be sure read Book «mentiond in» note 62. p.734.
     (How strange considering sweet Peas .)


lines 10—16 score in pale pencil
line 16 underline "gedenken"
line 16 annotation mention
line 17 underline "ausgezeichnete"
line 17 annotation marked out
lines 9—15 annotation Father & mother element more powerful in some
     ? do not understandt01
t01 - `? do ... understand' cancelled

lines 15—10 annotation Hybrid pollen more effect on on stigma than 2 other pollens; but the converse no effect

lines 5—3 score
lines 5—1 annotation another reverse case of different effects.


line 8 underline "Lychnicucubalus"
line 8 annotation Hybrid, I suppose
line 10 underline "Lychnis diurna"

lines 14—11 annotation colour & size of pollen no relation to fructification

lines 8—6 score
lines 8—4 annotation most important see his other work .

line 7 crossing-out "Varietäten"
bottom-margin annotation ⧟ (p. 181 of species «so holds good with species also») — From table at end, really speciest01
t01 - `— From ... species' in dark pencil
show subjects subjects

lines 7—5 double score
lines 9—5 annotation cancelled like Hollyocks, as if varieties
line 5 underline "fruchtbarer"
[illegible word]bottom-margin annotation ⧟ more fruitful , ∴ crossing cross colours less fruitful ∴ perhaps Hollyocks thus accounted for.t01 (So he says most distinctly in his Beitrage p. 137 in regard to Verbascum.)
t01 - `, ∴ ... for.' in dark pencil


lines 11—10 annotation (a)
bottom-margin annotation (a) Koëlreuter ‹mingled› «confused» ‹the› imperfect impregnation in th first cross, with th imperfect fructification of Hybrids, but this shows how similar th case is .—


line 7 underline in pale pencil "Grösse"
lines 9—11 score
line 10 underline "ohne ... worden"
lines 5—12 annotation imperfect fructification differs from no fruct., in seeds being more perfect

lines 17—20 underline "niemals ...angetroffen"
lines 17—22 annotation ‹hybrids› «first cross» never quite fertile as of cross of pure species.


line 8 score [`bookmark']


lines 10—12 annotation no relation in state of capsule & state of seed.

lines 17—27 annotation fertility of original act of hybridisation so different, that even in flowers of same plant, that it is difficult to make scale of fertility & consequent relationship.

bottom-margin annotation [illegible word]› Grades of Fructification, imperfect, so' normal & perfect «(a)» pollen no more effect than foreign dust; occurs even «not seldom» in species of same genus, “from want of sexual affinity”— (mere words)


lines 16—17 underline "er ... aus"
lines 15—16 annotation in brown ink Q
line 20 underline "schwaches ... Leben"
line 23 underline "1000"
line 25 underline "nicht ... gekeimt"
lines 21—25 annotation out of many imperfect seeds & ‹out of› 1000 seds apparently perfect, ‹of› «not» one germinated.


lines 1—2 annotation x
top-margin annotation ⧟ x every gradation of imperfection in capsule with one or more seeds, capalbe of germination.

lines 4—7 multiple score
line 4 underline "Abtheilung"
line 4 annotation division of subject
line 4 underline "selten"
line 5 underline "reinen"/ line 5 underline "Varietäten"
line 6 underline "Wahlverwandtschaft"
line 6 annotation affinity
line 7 at "Ergebniss" annotation result
lines 4—5 annotation Important

lines 12—15 score
line 12 at "littoralis" annotation ia
line 13 at "barbatus" annotation ia
line 13 at "diurna" annotation (ia & k)
line 14 at "mauritiana" annotation ia
line 14 at "glabra" annotation (k)
line 15 at "thapsiforme" annotation ia
line 15
lines 12—13 annotation in dark pencil most fertile hybrids

[continues overleaf] 5 underline "der freien Natur"


lines 2—6 score
lines 2—6
top-margin annotation in dark pencil Can the effect of pollen of plant in producing capsule be analogous to Ld Moreton's case?


line 10 score [`bookmark']


line 11 underline "Morton"
lines 12—17 score
lines 12—17 annotation Morton attributes power of hybridisation to capacity of domestication. —


line 9 underline "grosser"
line 11 underline "700"
lines 7—9 annotation greater number will not hybridise
line 11 underline "700"
line 12 underline "250"
line 12 at "Bastarde" annotation different sorts?

line 14 underline "versagt"
lines 13—11 score
lines 13—11 annotation unions which failed with Kölreuter


lines 15—14 underline "eine ... Elemente"
lines 17—9 annotation a certain sexual harmony necessary for union.— (I presume in contrast to general affinity)

line 4 at "haftet" annotation cleaves
lines 5—3 annotation pollen does not adhere to stigma


lines 8—15 annotation Even when pollen does cling to stigma frutification vry often fails
     In hybrids th stigma fails

lines 15—11 score in pale pencil
lines 10—9 underline "wie ... scheint"
lines 10—9 annotation (a)
lines 10—9 annotation in dark pencil only certain individuals can be hybridised
bottom-margin annotation (a) I cannot but think hybridisation commoner with animals than plants .


line 3 underline "angemessen"
line 3 annotation suitable


table underline "Primuleae"
table annotation easy
table annotation Fams. of easy manipulation

lines 26—24 score
line 21 underline "vermuthen "
line 10 annotation suspect ‹imagine›
lines 26—21 annotation Capacity for hybridisation not lies in Family Character.—

line 13 underline "Behuf"
line 13 annotation behalf
lines 18—11 annotation cancelled In families with regular specis , & divided into not real genera, most hybridisation

lines 9—8 underline "Gramineae ... zählen"
lines 11—5 score /lines 12—6 annotation The spec of most natural Families, viz .... resist hybridisation

line 2 underline "Compositen"
line 2bottom-margin annotation No hybrids in Compositae


line 15 underline "Apocineen"
lines 16—14 annotation ‹why› is not this Vinca
show subjects subjects
show subjects concepts

line 12 underline "unübersteiglichen"
line 12 annotation unsurmountable
lines 11—6 annotation thinks Orchideae & Asclepiadae wd be hard to cross., frm structure of flower


line 7 underline "Gymnogramma"

lines 16—19 annotation Disputes fern-hybrids, thinks only variations observed only in Gymnogramma


lines 16—20 annotation Dioecious less easy hybridised than hermaphrodite
show subjects subjects


lines 2—4 score
lines 3—5 score
top-margin annotation ⧟ The capability of hybrid‹isation› fructification ‹occur› lies in more special character, than those characterising any whole family

line 13 score [`bookmark']


lines 3—11 annotation No distinct relation between polyspermous & oligospermous plants & capacity for hybrid. fruct:.—

line 7 at "Vermeidung" annotation avoidance


lines 1—2 score
line 1 underline "eine ... Blüthe"
line 2 underline "Conceptionskraft"
top-margin annotation Dioecious plants a longer capacity for impregnation .    I see uses Dichogamous = Dioecious

line 5 underline "Lecoq"
lines 3—5 annotation a book on Hybrids }—
show subjects concepts

lines 11—9 score
line 10 annotation (a)
bottom-margin annotation a    Dioecious plants less capalbe of Hybrid-fruct: at least than some hermaphrodites


line 7 score [`bookmark']

table at "Anagallis; Potentilla; Salvia; Cucumis; Lycium" annotation
table annotation ⧟ other observers have succeeded, though he failed.


line 8 at "fügen" annotation join
line 9 at "zuverlässigen" annotation certain

line 10bottom-margin annotation closely allied genera differ greatly in tendency to hybrid-frutification several examples & I believe quotes Kolreuter but observes only few species in each experimented on . —

lines 6—5 underline "Aquilegia"
lines 6—3 annotation cancelled Hooker thinks all same species
show subjects concepts


line 2nd table score

line 16 at "ununterscheidbare" annotation undistinguishable
line 18 underline in dark pencil "Afterbefruchtungen"

lines 21—22 underline "Afterbefruchtungen irregeführt"
line 22 annotation lead astray
line 23 at "wähnten" annotation believe
line 3rd table score
line 20bottom-margin annotation All this shows that when anther removed, how much crossing can take place from adjoining plants — i.e. intermarriage


line 12 at "Einreden" annotation objections
line 12 at "veranlasst" annotation caused —


lines 10—9 score
lines 11—6 annotation I do not know whether Kolreuter or self. he suceeded anyhow.

lines 10—7
lines 7—3 score
lines 5—1
line 5 underline "vespertina ... Cucubalus"
line 4 at "noctiflora" annotation (yes) ,
line 1 underline "wiederholten Versuchen"
lines 7—5 annotation Bigeneric crosses


lines 1—2 score
lines 1—3 score
line 1 at "Kreuzung" annotation reverse
line 2 underline "♀"
line 2 underline "♂"
line 3 underline "vergeblich ... hatte"
line 2 annotation (a)
top-margin annotation (a) Revrse case which always failed with Kolreuter succeeded once with him,— but was vry difficult Hybrid Plants no ways different —t01
t01 - `but was ... —' in dark pencil

line 3 crossing-out "8"
line 3 annotation 107 /
line 14 crossing-out "9"
line 14 annotation 108 /
show subjects concepts


line 8 score [`bookmark']


lines 11—14 score
lines 11—17 annotation universal law that pollen of parents, fructifies hybrids more then own.

line 20 annotation / 123
show subjects concepts


lines 4—1 score
lines 4—2 score
lines 3—2 underline "Canis ... Mouflon"
line 1 underline "zahme ... Mouflon"
line 1 at "Böcke" annotation ⧟ ram or he goat


line 3 crossing-out "8"
line 3 annotation 137
show subjects concepts


lines 8—22 score
lines 12—15 annotation (a) → sense given abovet01
top-marginline 7 annotation in dark pencil (a) Against ‹law› «limit» of genus being determined by «power of» crossing , even Herbert does not pretend all species can cross, though when any true species do cross, he says thy must belong to same genus — so th “reverse crosses.” & cases of Crosses which after years succeed only once, go against law of genus & crossing being connected.
t01 - `sense given above' in dark pencil

[continues overleaf] 3a "in" annotation power of uniting depends on
lines 3—2 underline "inneren ... Arten"


[continuation] 1—2 underline "systematische"
line 2 underline "sexuelle Verwandtschaft"
top-marginline 1 annotation Hence a sexual & systematic relationship

line 15 annotation in dark pencil 138 /
show subjects concepts

lines 17—20 annotation genera formed of diverse ‹ty› characters


lines 6—16 score in pale pencil
lines 6—11 → in pale pencil
lines 1—3 score
top-margin annotation When two genera have united, th species do not always in these genera readily unite. Tormentilla & Potentilla , though so close ,will not unite
from Slip Attached 1, Side 2 annotation in dark pencil Did Kolreuter cross many Silene «vide» p. 140 of Gaertner . ? /

line 1 crossing-out "141"
line 1 annotation 0
show subjects concepts

lines 17—30 annotation / it is quite wrong to infer becaus some species within a genus, will cross that all will; generally only a few. Herberts cases, however, rather contradict this . Hippeastr, Azalea, Calceolaria


lines 18—10 score in pale pencil
lines 16—14 annotation Sections of genera
lines 6—5 score
lines 6—5 underline "blauen ... gelben"
line 1 underline "Zephyranthes"
lines 1—6 score
line 3 annotation (a)
top-margin annotation (a) Sections of genera will sometimes unite & sometimes not.—

line 8 crossing-out "2"
line 8 annotation 1
line 6 crossing-out "3"
line 6 annotation cancelled 2
show subjects concepts


lines 8—11 annotation in dark pencil in his Treatise speaks of species see to this
lines 11—13 score
line 12 underline in dark pencil "Varietäten"
line 12 annotation deleted in dark pencil species?t01 genera of
lines 10—12 annotation ??
line 13 at "Pepo" annotation pumpkin
line 14 at "Citrullus" annotation water melon
line 15 at "wiewohl" annotation though
lines 15—16 annotation in dark pencil Girou succeeded in these.
from End Note annotation p. 142    See to this important
t01 - `species?' in dark pencil

line 13 crossing-out "6"
line 13 annotation 5
line 15 crossing-out "7"
line 15 annotation 6
show subjects concepts

line 19 at "Verzeichnissen"
line 19 annotation in dark pencil statements
lines 20—28 annotation The list of sections of genera, which will not cross, shows no necessary relation of genera & crossing.—


line 6 underline "sexuelle Affinität"
lines 6—11 annotation sexual affinity belongs only to species, & often to only individuals, & cannot be externally recognised.
show subjects concepts

lines 12—16 annotation Different Times of flowering do not prevent hybrids.

lines 3—2 underline deleted "Lavatera ... trimestris"
bottom-margin annotation ‹Bush with plant ?›


line 1 at "strauchartige" annotation bush    with
line 1 at "krautartigen" annotation herb

lines 3—6 annotation annual, biennial & perennial cross.
show subjects concepts

lines 16—19 annotation evergreen with deciduous cross
show subjects concepts

line 17 at "erweckt" annotation awaked

lines 16—7 annotation many hybris wh suceed in summer, & fail in Autumn — some succeed so seldom may be called accidental.

lines 2—1 double score
bottom-margin annotation the most different varieties can cross.—


line 3 underline "Phaseolus, Pisum"
line 3 annotation Can cross

line 14 underline "auch ... bestätigt"
line 14 annotation see to this


line 17 underline "obgewalten" annotation exist


line 10 underline "Umfang" annotation extent

lines 12—10 underline "Nicotiana ... glauca"
line 10 underline "die rustica"
lines 15—7 score
lines 15—7 annotation Pollen of some plants can impregnate others, but not be impregnated. by them.

lines 7—1 score in pale pencil
line 6bottom-margin annotation case of Nicotiana which will neither impregnate or be impregnated by other close species.
bottom-margin annotation in brown ink 8 specs


[continuation] 1 at "diese" annotation varieties
top-margin annotation Van Mons thinks stability in first generation & variability in first distinctive character of species .

lines 8—13 double score
lines 7—11 annotation analogous to some organisms not varying in domestication

lines 8—9 underline "Dianthus ... Crinum"
line 10 underline "sechste ... Generation"
lines 9—15 annotation some hybrids as these keep constant in 7th & 8th genertns, but this rare.—

line 14 underline "willkürliche" annotation
line 14 annotation arbitrary

line 12 underline "E. Blyth (157)"
line 12 annotation see to this
line 10 at "anderer" annotation of anoth kind
line 9 at "jene" annotation species

line 3 at "Eichelheher" annotation Jay

bottom-margin annotation I had better give cases of closely allied «& identical» species in different climates to show species , & vry different species in similar climates to show not direct effect of climate
show subjects subjects


line 1 underline "unbedeutenden" annotation insignificant

top-marginline 15 annotation I quite agree vry near or identical species may have been created , but this has to be decided.

line 14 score
line 14 underline "vergesse"
line 14 annotation forgets

lines 17—22 score
line 20 at "Thurmschwalbe" annotation swift ?
line 19 annotation (a)
bottom-margin annotation (a) A great power of place-alteration has no relation to wide geographical range. — in direct opposition to my view, but agres with Hooker on Compositae.

lines 20—30 annotation Seems to agree with Blyth,, I must study him —


line 9 at "Abkunft" annotation [illegible word]
line 9 at "vereinigt" annotation [?]fixed
line 12 at "eingewendet" annotation opposed


line 5 underline "aufgehört" annotation ceased
line 6 underline "Stillstand" annotation situation
line 10 at "abgeschlossene" annotation settled

line 1 underline "Rosa ... Nicotiana"
line 1 annotation genera with mny doubtful species


line 2 at "anerkannt" annotation acknowledged
line 6 at "stete" annotation fixed
line 7 at "eingeprägte" annotation inculcated
line 6 underline "vertilgbar" annotation exterminate


line 18 at "Lücke" annotation gap
line 5 at "Huldigen" annotation do homage


line 12 at "Beurtheilung" annotation judgmnt
line 11 at "Vertheidiger" annotation defender


line 15 at "naturgemässen Verhältinssen" annotation conformable to nature    relations


line 1 at "Gegenstand" annotation resistance
line 8 at "Anspruch" annotation lay claim to


line 17 at "vorgeblichen" annotation pretended
line 9 at "Ermittelung" annotation enquiry


lines 9—11 score in pale pencil
line 10 at "Bedenklichkeit" annotation doubtfulness
lines 6—11 annotation I think it means only that result of A.B.C with ξ differ greatly from each other
     (a)

bottom-margin annotation (a) Is there any case of two «close» vars , when united to a 3d «var.» producing vry different mongrels.

line 15 underline in dark pencil "nahe verwandter"
lines 15—22 score in dark pencil
lines 18—19 underline "z. ... fulgens"
lines 21—22 underline in dark pencil "geben ... Bastarde"
lines 12—19 annotation in dark pencil, cancelled in pencil hybrids from close species when united with one another, show their differences even plainer than whilst pure. —

lines 15—8 score
lines 16—10 annotation Remarks This very odd if these close species descended from common ancestor.

lines 8—3 score
lines 16—1 annotation in dark pencil The existence of species consists of fixed sexual relation to other species


line 11 underline "streitig"
line 11 annotation disputable
line 16 at "Gültigkeit" annotation valid

lines 17—18 underline "die ... fruchtbar"
lines 16—20 annotation in dark pencil Koelreuter's law of sterility can hardly be accepted.

line 13 underline "Pentstemon ... coccineus"
lines 14—12 annotation do thy seed?
line 4 annotation X
lines 14—6 annotation in dark pencil These 2 Penstemons though so like, as to be considered varieties, cannot be crossed.

lines 3—2 score
bottom-margin annotation ☞ ||    rarely certain individuals will not be impregnated see G. Beitrage.


line 5 at "krankhafter" annotation sickness

lines 17—13 score in pale pencil

bottom-margin annotation not in Darwin's hand X There are two P. gentianoides in cultivation the one commonly so called is the Hartwegi so misnamed — the true gentianoides is rare in cultivation differently shaped and not red


line 9 underline "der ... seie"
lines 8—12 score


lines 15—10 score
lines 16—13 annotation nearly related but will not cross

line 4 underline "H. Lecoq"


line 3 at "Veranlassung" annotation cause
top-marginline 5 annotation I fancy that this is only that parents have originally crossed.

line 12 "Varietätenbastarde"
line 13 at "verhalten" annotation are circumstanced
lines 8—17 annotation I do not understand.t01 { are «circumstanced» ‹like› «as» th second generations of species-bastard    Will explain more afterwards
t01 - `I do ... understand.' cancelled

lines 17—22 score
lines 14—24 annotation Holds good with some wild ‹variety› ‹variations› «species» which fructify each other but do not sport like true vars, whereby these plants are characterised like true species.

lines 19—26 score
line 21 underline in brown ink "beinahe vollständig"
lines 19—24 annotation Get information on these — wild ‹vars.› «species» Fertility tested by himself.

lines 7—6 score
line 6 annotation a
bottom-margin annotation (a) Genera with species agreeing in Habit, as above, hybridise most, —yet some species of these will not cross.


lines 4—6 score in pale pencil
lines 4—7 score
lines 3—7 annotation These species cross easily, yet other others of the genus ,will not cross.

lines 14—15 underline "Gramineae ... Leguminosae"
line 16 underline "Geranium ... Myrtus"
lines 9—17 annotation Though power of crossing sometimes goes with external resemblance yet the most natural Families & genera as here, do not hybridise. well.—

line 13 at "zuverlässige" annotation positive
line 12 score
lines 14—13 annotation Hybridisation difficult
show subjects subjects


lines 1—4 score
lines 1—4 annotation Most Natural Family
show subjects subjects

lines 9—14 score
lines 9—10 annotation all failed

lines 11—9 score
line 10 underline "Sageret"
line 10 underline "Lecoq"
lines 12—10 annotation quotes from Sageret
line 9 underline "ohne Unterschied"
line 9 annotation promiscuously
lines 11—1 annotation Sageret & Lecoq has found vars of Cucurb. will not cross promiscuously (References hardly bear out conclusion)


lines 1—7 score
top-margin annotation cases of only few species in very close genera uniting, even species «which are» hard to specifically characterise.—

table annotation in dark pencil cases of very close species ‹which› or more exactly, species having th same habitus which will not unite.—

[continues overleaf] lines 5—1 score
line 5 underline "Lychnis ... cuculi"
bottom-margin annotation cases of species having very different habitus which do unite, chiefly from Herbert, except Derby


[continuation] 1—8 score
line 9 at "transsylvanica" annotation Köelreuter

lines 14—10 score
lines 14—8 annotation So Koelreuter shows, that propinquity does not go with power of hybridising
lines 7—1 score
lines 7—1 annotation other examples of the same law


line 2 at "Langsdorfii"
top-margin annotation cannot be fructified by the 4 named sorts, though it can fructify them & some easily.

lines 10—20 score
lines 10—20 annotation even when mutual crossing does take place in closely allied species, yet facility not alike (this is new)

line 21 underline "Tropaeolum ... minus"
lines 21—22 score
lines 21—27 annotation closely allied, yet unite with difficulty & will not be reciprocal.


top-margin annotation horizontally crossed || officinalis & acaulis not in Loudon.— p 721 officinalis = veris = Cowslip — I see it is barely possible without consultng Babington to know which is which
from Slip Attached 2, Side 1 annotation      (For p. 178)t01
     1. P. acaulis P. vulgaris.    Primrose. a speciest02
     2 P. elatior (calycantha)
     3 P. elatior (communis)    — — P. elatior.    Oxlip, true plant. a speciest03
     4 P. veris (officinalis)    — — P. veris.    Cowslip. — — a speciest04
     5 P. veris (officinalis) calycantha
     6 P. calycantha (elatior)
     N.B. in Nr 3. “communis” sometime in Bracket, sometimes not.—
        No 5. I presume must be some blunder of Printers
        No 6. ■ Can these names have been reversed by Printer
t01 - `(For p. 178)' in pencil
t02 - `P. vulgaris ... species' not in Darwin's hand
t03 - `— — P. elatior ... species' not in Darwin's hand
t04 - `— — P. veris ... species' not in Darwin's hand
show subjects concepts

line 3 underline "officinalis"
line 3 annotation cowslip
line 3 at "elatior" annotation oxlip
line 4 underline "acaulis"
line 4 annotation primrose
top-marginline 3 annotation x----(Q)
top-marginline 8 annotation /Most important
     on Babington's authority see Table

top-margin annotation Here are vars ,which will not unite
from Slip Attached 2, Side 1 annotation      (For p. 178)t01
     1. P. acaulis P. vulgaris.    Primrose. a speciest02
     2 P. elatior (calycantha)
     3 P. elatior (communis)    — — P. elatior.    Oxlip, true plant. a speciest03
     4 P. veris (officinalis)    — — P. veris.    Cowslip. — — a speciest04
     5 P. veris (officinalis) calycantha
     6 P. calycantha (elatior)
     N.B. in Nr 3. “communis” sometime in Bracket, sometimes not.—
        No 5. I presume must be some blunder of Printers
        No 6. ■ Can these names have been reversed by Printer
t01 - `(For p. 178)' in pencil
t02 - `P. vulgaris ... species' not in Darwin's hand
t03 - `— — P. elatior ... species' not in Darwin's hand
t04 - `— — P. veris ... species' not in Darwin's hand

lines 4—9 score
lines 4—8 annotation cancelled Converse of these very difficult

lines 10—14 score in dark pencil
lines 10—12 score
lines 13—14 score
lines 11—16 annotation in dark pencil very different in Habit, yet unite easily.—


lines 7—12 score
line 10 underline "Cereus"
line 11 underline "schon längst"
line 11 underline "Cereus"
line 8 annotation Herbert

lines 13—19 annotation Cactus or Cereus Melocactus Echinocac, Echinos, Phylocactus
line 20 underline "H. Neubert"
lines 21—22 underline "Cereus ... Ottonis"
line 24 underline "flagelliformis"
lines 24—25 underline "speciosissimus ... phyllanthus"
line 25 underline "grandifloris"
lines 27—28 underline "Echinocactus ... Phlylocactus"
lines 24—28 annotation Neubert has succeeded in these crosses


lines 3—6 score
lines 2—7 annotation Mongrels sport, & he has seen same thing Cucurb:
show subjects subjects

lines 18—15 score
show subjects concepts

lines 12—8 annotation in dark pencil Flowers vry unlike yet cross

[continues overleaf] lines 3—1 score in dark pencil
lines 5—1 annotation in dark pencil Colours of Verbscm


lines 6—5 score [`bookmark']


line 7 at "grellen " annotation distinct
line 6 at "enthalten" annotation contain
line 2 at "beiträgt" annotation contribute


lines 6—8 score
top-marginline 10 annotation cause of Hybridising a vital action & allows that the sexual relations is mere word

line 15 at "Leitfaden" annotation leading-line

[continues overleaf] lines 16—15 double score
lines 17—16 underline "sondern ... beide"
lines 12—1 score in pale pencil
lines 22—3 annotation Summary but nothing new


line 10 at "giftige" annotation poison
lines 6—14 score
lines 6—14 annotation cases of plants differing chemically , compares this with difference in their sexual affinity .

line 15 at "Wahlverwandtschaft" annotation in dark pencil "elective affinity"


lines 3—13 score
line 4 at "Begriff" annotation idea
line 7 underline "Wahlverwandtschaft"
line 7 annotation “elective affinity”
top-margin annotation Fertility of Hybrid , ‹depe› is in even less reation [ie `relation'] to affinity of parents than facility of first ‹concours› «union or hybrid-fructification.» . It seems no relation between ease of getting 1st hybrid & this hybrids fertility . This is case with the common Mule.—
show subjects subjects

lines 11—16 score
line 11 underline "manche"
line 12 underline "leicht"
line 16 underline "sexuell ... verwandt"
line 16 underline "49"
line 18 underline " waren ... fruchtbar"
lines 9—16 annotation in dark pencil many plants easily cross, whose hybrids are quite sterile

line 18 at "Kölreuter" annotation (a)

lines 14—13 score
lines 15—10 annotation and fertility of similar Hybrids very variable

line 12 at "Wahlverwandtschaft" annotation ‹sexual› elective affin
table annotation Hyb. fruct. of great difficulty
bottom-margin annotation I think Verbascum is case in point.


line 15 underline "80 ... 120"
lines 17—14 annotation seeds in pure ‹[illegible word]› parent cross
show subjects subjects

line 12 underline "paar ... Samen" /lines 13—9 annotation numbers of seed ‹in [illegible words]› «Hyb. fruct:» when crossed.
show subjects subjects

line 10 underline "151"
line 9 underline "29"
bottom-margin annotation ⧟ numbers of seeds in reciprocalt01 Hyb. frucctification.
t01 - underlined in brown ink

bottom-margin annotation (hybrd-fruct. best expression)
     (pure. fructification)


top-margin annotation ☞ (Bad simile) We might as well as deny that th different «substances»t01 were really different, because thy had no “elective affinity” «(I use word of Gärtner)» t02 whereas other two had strng elective affinity & wd. unite & make a third. —
t01 - `substances' in dark pencil
t02 - `« (I use ... Gärtner)» ' in dark pencil

line 4 annotation («sexual» non-reciprocity) of th “elective-affinity”.—


lines 1—4 score
lines 6—8 score
top-margin annotation The reciprocity of sexual alliance is not only different in strength, but is often entirely deficient

lines 12—24 score
lines 12—24 annotation cases of slight unequal reciprocity in very closely allied speces, some even thought to be varieties.

lines 4—1 score
lines 4—1 annotation cases of more unequal reciprcity


lines 8—12 score in pale pencil
lines 7—9 annotation (z)→
top-margin annotation (z) || The absence of perfect reciprocity even in nearly related species, shows that male & female power of union do not go together : but this difference of male & female has no effect on th Hybrid offspring


lines 8—12 score
lines 10—11 annotation (a)
lines 7—5 annotation do →t01
top-margin annotation (a) N. rustica will not unite with N. glutinosa, but hybrid N. paniculata—rustica, will with glutinosa, & the character of rustica is seen in offspring:—
t01 - `→' in pale pencil


lines 1—13 annotation as in Primula & Verbascum (?) But then question of what species are is begged —
show subjects subjects

line 15 underline "Kölreuter"
line 16 underline "hybride Befruchtung"
lines 14—17 annotation ‹Hybrid-fruct› «First cross» never bring so many seeds as nat. fruct.

line 4 underline "jedem"
line 4 annotation frm each
lines 2—1 score
bottom-margin annotation Remarks that Herbert's Crinum was not growing in own climate — But he says evry ovule was impregnated.


lines 11—18 score
lines 14—15 annotation (a)
top-margin annotation (a)    Can judge of scale of elective affinity by number of seeds in hybrid-fruct—, as compared to normal fruct.

line 16 underline "Schätzung" annotation valuing

lines 13—1 score
line 10 underline "nicht constant"
lines 9—8 underline "doch ... gross"
lines 2—1 underline "Mittel ... entwerfen"
bottom-margin annotation an average of seeds taken from a number good plant growing in open nature


line 7 at "Vorschlag" annotation calculation
line 9 at "beabsichtigen" annotation have in view
lines 10—12 score

lines 11—6 score
bottom-margin annotation on account of unfavourable conditions , we take maximum of seed of hybrid cross .


line 2 X [`bookmark']

lines 13—22 score
lines 13—22 annotation in wild Plants nmber of seeds do not differ so greatly as to cause much difficulty in estimatng numbers.

lines 4—2 ]
line 6bottom-margin annotation There are ‹are› differncs in flowers of same plant in being impregnated by foreign pollen.—


lines 8—10 score
lines 8—9 underline "dass öfters ... also"
lines 2—8 annotation as individuals differ in som respect, as last page, several must be experimented on—

lines 16—19 score
lines 16—19 annotation cases of individual plants wh. were femally sterile

line 14bottom-margin annotation As th difference between nature & artificial self-impregnation is never so great as in Hybrid fructification; he has taken wild plants as base of calculation, which seems to me to be an error.

lines 4—3 underline "selbst ... käme"
bottom-margin annotation Effects of crossing every plant by self injurious.— see Beitrage p.366


line 14 double score [`bookmark']

line 8 at "vermeiden" annotation avoid


lines 2—9 annotation Sexual affinity calculated by maximum of good seeds, till further experiment ever increase this maximum.

lines 7—1 score
bottom-margin annotation experiments shd. be tried at different times on different plants.


lines 5—7 score
line 5 underline "vollkommensten"
line 5 annotation (a)
line 14 underline "keimungsfähigen"
top-margin annotation (a) Take average of number of seeds «capable of germination» under normal circmstances as the standard for comparison of best fruits

lines 14—12 apparently unintentional mark in brown ink

lines 6—1 score
line 4 underline "20 Versuchen"
bottom-margin annotation in dark pencil Very important, if this smaller number be not due to ‹Art› «art» used in the fructification.


table at "polline" annotation naturally ‹f› impregnated

table annotation in dark pencil Scale of sexual elective affinity, inferred frm maximum seeds from hybrid-fruct, not from Hybrids themselves.
show subjects subjects

table underline "0,6224"
show subjects concepts


table underline "0,6666"
show subjects concepts

lines 9—3 score
lines 8—7 annotation (a)
bottom-margin annotation (a) «Gärtner» thinks that these tables of affinity show that pure species are aboriginally formed stabile.— It is contradicted absolutely by his vars.t01
t01 - `It is ... vars.' in pale pencil


line 1 at "Cucubalus" annotation ⧟ Here a genus more fertile than other species    777/1000
line 1 at "Cucubalus" annotation Silene of Steudel /


top-margin annotation cancelled 0011/10000
show subjects concepts

table 18—21 score
table 18—21 annotation 3 genera before other species


line 4 underline "Kreuzung"
line 4 annotation with G. & Kolreuter implies reciprocal fertilisation

lines 3—1 score
bottom-margin annotation cases of non-reciprocal frutification .


top-marginline 24 annotation in dark pencil self & Köelr. find these reciprocal crosses exactly alike.
line 19 underline "allgemeine"
lines 19—24 annotation This is general rule, specially in wild plants, which are not varieties.

lines 22—23 underline "Abweichungen ... Farbe"
line 26 underline "Ausnahmstypen"

line 10bottom-margin annotation Difference frm Animals as Mule & Hinny . also hybrid animals differ in sam litter; but in animals all half domesticated.
show subjects subjects


lines 4—7 unmarked
lines 16—14 score
line 14 underline "Differenz im Habitus"
top-marginline 5 annotation in dark pencil Diff in animals & Plants owing to sexes separate in animals.

line 10 score [`bookmark']

lines 8—3 double score
lines 7—6 annotation Hybrds varid

lines 25—2 annotation In comparison of sexes we must suppose habitus the same, & form of parts direct result of sexual peculiarities. Whiskers in Man !!t01
t01 - `parts direct ... !!' in dark pencil

bottom-margin annotation in pale pencil Whiskers & Mane cannot be thus accounted for.

bottom-margin annotation No difference in Habitus of Plants, when sexes separate, (because I say do not struggle for female: so lower radiata.
show subjects subjects


lines 17—20 annotation slight variations in hybrids


line 14 score [`bookmark']


lines 16—12 score
lines 17—10 annotation Double flower raised from male or female


lines 4—9 score
line 4 annotation a
top-margin annotation Differs from animals for sex no effect on Hybrids
lines 11—14 score
lines 12—13 annotation see to this


lines 14—18 score
lines 20—24 score
lines 12—24 annotation Form of hybrids stable in 1st generation → does not hold good with animals.


lines 8—10 score
lines 8—10 annotation experimentised with wild plants


line 18 underline "in der ersten"
line 10bottom-margin annotation It is proved by long course of his & Köl's exprimnts that bastard even «in» 1st. generation frm same parents are always alike; & return in constant course to either parent, when repeatedly crossed with such. → (a) Thinks this evidence of permanence of species; but I do not see more than ordinary generation keeping true; perhaps tests the trueness in another way: but a plant does not vary in first generation, when put out of normal conditions.

lines 13—5 score
lines 24—10 annotation (a) Notice this argumnt


lines 5—7 score
line 6 at "Entstehung" annotation origin
line 6 annotation a
top-margin annotation (a) Hybrids unions therefore follow same law in first generation as the union of pure species.—

line 8 score [`bookmark']

lines 19—21 score
lines 19—21 underline "So ... erhalten"
lines 9—21 annotation Thinks the few exceptions to the normal structure of hybrids is due to variation (a)
bottom-margin annotation (a) Rather hard, it seems to me to draw distinction; but Gaertner (z) urges the resemblance of Hybrids made at same time & after long intervals from same pure parents.

lines 12—5 score
lines 10—9 annotation (z)


lines 10—11 score
lines 8—14 annotation The normal Hybrid type keeps constant in succeeding generations only in the most fertile hybrids, generally.


line 1 at "Ausnahmstypen" annotation Exceptional

line 9 underline "einzelne ... Bildung"
line 11 underline " sehr ... einem"
line 12 underline "doch ... mehreren"
lines 6—14 annotation very rarely sometimes single sports in a set of normal hybrids out of same fruit; &


line 2 score
line 2 underline "Digitalis, Lobelia"
top-margin annotation only genera in which these exceptional types have been observed: (z) These exceptional types from same species always resemble each-other.!!

lines 12—15 score
lines 11—14 annotation (z) →

lines 12—7 score
line 13bottom-margin annotation on two years a peculiar yellow var, so unlike as might be thought a different kind.— The mother type prevailing


lines 5—8 annotation in dark pencil one specimn of abnormal type.

bottom-margin annotation I observe that these abnormal types often take after one parent.


line 11 score [`bookmark']

lines 16—23 annotation ☞ abnormal types generally quite sterile; (this very curious)

line 7 at "Impfung" annotation inoculation
lines 9—4 annotation compares these abnormal types to atavism

line 4 at "Gleichgewicht" annotation equilibrium


line 2 at "wankend" annotation staggering


lines 10—7 score
lines 9—7 double score
lines 9—7 underline "Mathiola ... officinalis"
lines 16—6 annotation (a)    These varieties seldom repeat each other.
     They occur chiefly in such species as are so closely allied, as to be held mere varieties

bottom-margin annotation The irregularity of reappearance, & slightness of differences, seem only distinct differences with his abnormal Hybrids of the previous chapters.—


lines 12—16 double score
line 13 annotation (a)
bottom-margin annotation (a) The abnormalities in Hybrids has observed only in plants, long cultivated in gardens, & not in wild plants; but I remember that only one side wild in Kölreuter is sufficient


line 11 at "fügt" annotation adds
line 6 underline "Räthselhaftes" annotation emigma


lines 20—25 annotation ‹generally› «not seldom» in Hybrids one side «or species—» prevails over other; & this prevailing is not accidental but is constant.

line 3 score [`bookmark']


lines 4—1 score in pale pencil
lines 3—1
bottom-margin annotation most difficult which of two parent-forms a hybrid comes nearest to


lines 3—7 score in dark pencil
top-margin annotation in dark pencil Does not believe that Hybrids are ever unlike both parents

line 20 score [`bookmark']


line 1 at "indem" annotation whilst
lines 5—7 score
line 6 annotation (a)
top-margin annotation (a) When two hybrids «‹but not when two pure plants›» unite, & ‹one› «offsprng» takes almost exclusively after ones side, hybrid is sterile.

line 11 underline "Lychnis flos cuculi"
lines 11—12 underline "Cucubalus behen"
lines 14—18 score
lines 14—18 annotation cancelled one specn took most closely after father ; was fertile.


lines 14—1 annotation odd that this hybrid no rudiment of stamen, considrng structure of both parents.

line 9 score [`bookmark']

lines 5—3 score
line 4 annotation B
bottom-margin annotation (B—) Female organ «generally» shows no signs of imperfection even when perfectly sterile.


line 9 underline "Thiervarietäten"

lines 19—21 score

bottom-margin annotation The entire differences, of different authors in ascribing more or less to Father or Mother shows that no real rule.


lines 14—11 score
lines 14—13 annotation “    ”
line 12 underline "pater major"
lines 14—11 annotation seems pretty true


line 12 score [`bookmark']


line 3 score [`bookmark']

line 5 underline "oben ... 202"

lines 1—24 annotation See in Kolreuter whether vars. with a species give vry similar Hybrids —

line 19 at "Specifische" annotation (a)
lines 19—20 underline "Specifische ... reinen"
line 20 underline "deutlicher"
bottom-margin annotation (a) [On Datura see my Abstract of Kolreuter p. 8]Bis]
     I see no reason why varieties shd not equally show this distinctness [corrected from `distinction'] in same way . Does not Ancon Sheep impress offsprng vry remarkably ??

line 23 underline "Stramonium ... Tatula"
line 26 underline "ganz ... Bastarde"
lines 22—26 score
lines 22—26 annotation different species because Hybrids different

lines 11—9 underline "magnifolia ... pumila"
line 11 at "macrophylla" annotation Loudon mak distinct
line 9 at "asiatica" annotation and these vars. of rustica
line 9 at "humilis" annotation not in Loudon
lines 12—8 score
lines 12—8 annotation These with N. glutinosa give ‹vry› «quite» similar product & therefore considers them vars.
     / accordingly th several vars of dogs ought to produce similar Hybrids with wolf

lines 3—2 double score
lines 3—2 annotation I do not understand


lines 11—15 annotation External conditions no special influence on character of Hybrids.

line 12 score [`bookmark']


lines 4—8 annotation Hybrid Dianthus more stabile than other genera —

line 12 underline in pale pencil "Caryophyllus ... simpl"
lines 10—12 annotation Varieties› Mongrels follow different laws to Hybrids /

lines 13—16 annotation Digitalis .peculiar. in its sporting, & exceptional.

lines 17—15 score
lines 17—15 annotation G
bottom-margin annotation G Thinks the by far greater number of normal to abnormal Hybrid types opposed to their resulting from external circumstances.

lines 7—6 score
lines 8—6 annotation In embryo-plant no alteration in Hybrid from mother
     embryo of mother


lines 1—3 unmarked
top-margin annotation In Hybrids, form of the cotyledons affected


line 11 at "auszuweichen" annotation evade
lines 12—19 score
line 15 annotation differ

line 15 annotation gemischte

whole-margin annotation I think children of pure parents, thus go after either one or other or intermediate but as he says for Hybrids ‹are› «how» difficult to judge & compare


lines 2—4 score
line 7 score
top-margin annotation In the cross Lychnis quite prevails over Silene, so that Hybrid is like var. of the Lychnis


line 3 annotation sterile
show subjects concepts

line 14 O
show subjects concepts

lines 20—21 underline "grandiflora ... wird"
line 18 annotation sterile

line 15 O
line 13 O
lines 10—9 O
lines 7—6 O


line 14 at "Decidirte" annotation When a hybrid most strongly resembles either parent

line 14 annotation Puts strongly how these 3 classes blend into each other..

line 1 underline "decidirt ... mütterlich"
bottom-margin annotation ‹See in› These terms used when th two have been crossed not reciprocally — Relativ-vaterlich &c used, when thy do cross reciprocally & take after th father in cross specified.—


lines 10—15 score in pale pencil
lines 9—17 annotation No relation ‹of› «in» closer resemblance to Father or mother to fertility of Hybrid.

lines 15—17 score
lines 15—23 annotation reciprocal crosses take place ‹when› «thogh» the offsprng take decidedly after one parent.

line 20 at "syphilitica" annotation } Decided types generally sterile,— but not always as in Lobelia

line 4 annotation strongest example

lines 3—1 score
bottom-margin annotation ‹What are› These Hybrids , ‹I presume› «are a» cross of Hybrid ‹& pure, &› «& a» third species — How is it «called» when two Hybrids cross ?t01
t01 - `How is ... ?' cancelled


line 4 score
line 4 annotation a
top-margin annotation Perhaps gemengte (or 2d class) the commonest p. 282.—

line 9 score
lines 8—7 annotation (a)
bottom-margin annotation (a) So there are species of genus which a prepotent ‹fructifyng› «fertilsng» power on other species ; so others have predominatng influence on structure of Hybrids; but these two are not connected


lines 1—2 score
top-margin annotation Even in th mixed types , one part now resembles one parent , now another part the other parent. —


lines 10—17 score in pale pencil
line 10 annotation (a)
top-margin annotation No relation between facility with which A will impregnate B or be [corrected from `being'] impregnated by it, in different cases, & th resemblance of hybrid to th parent.—

[continues overleaf] lines 3—1 score
bottom-margin annotation This seems a Lamarkian
from End Note annotation Treviranus «p 292» ought to be read .    He seems a Lamarckian.


lines 1—3 score
line 2 annotation a
lines 7—9 score
line 8 annotation a
top-margin annotation ‹The› Always something new in appearance of Hybrids; but not absolutely new, but appear so from odd unions & opposition of parent forms .

line 17 underline "Mirabilis"
line 17 annotation } cases of in
lines 17—20 annotation N.B Both sporting genera


line 19 score
lines 17—12 annotation Colour variable often, in crosses & unexpected


lines 13—18 score
lines 13—20 annotation flowers do not take after Mother or Father in colour.—

[continues overleaf] 2 underline "Dianthus"
line 5bottom-margin annotation Some simple Hybrids retain in successive generations their colour, as in Dianthus &c.
     But generally (next Page) case very different; colour most variable


[continuation] 2—5 score
lines 2—5
line 3 crossing-out "Aleca"
line 3 annotation Hollyoc
lines 3—4 underline "Vinca ... coerulea"
top-margin annotation variation said to keep true.— Vinca rosea a stove Plant. Syring Lilac. p 743. The Book quoted probably cd not be consulted

lines 8—14 score
lines 8—14 annotation very variable colours in successive generations of Hybrids

lines 19—24 annotation complex Hybrids even more variable in colour.

line 10 underline "der zusammengesetzten"
lines 10—4 annotation ⧟ These hybrids take almost always colour of father .


lines 1—2 underline "Lychnis diurna einen"
lines 4—5 underline "Mirabilis ... centifolia"
line 8 underline "drei verschiedenen"
top-margin annotation Sports

line 14 underline "Achillea Millefolium"
line 14 annotation Sport
show subjects concepts

lines 12—7 apparently unintentional mark


line 10 score
lines 9—7 score
line 8 annotation (a)
bottom-margin annotation White flowers commoner here than more South.—


lines 1—4 score
line 2 annotation (a)
top-margin annotation Important on account of ‹Verbascum› Kolreuter . Verbascum Lychnitis with white flowers ,rarely with yellow on sandy Places —
     ☞ (So Kolreuters case goes for nothing)

lines 8—12 score
lines 8—10 annotation seed from yellow gave chiefly white

lines 14—20 score
lines 12—19 annotation when crossed colours did not mix, but came pure yellow or white
show subjects subjects

lines 6—20 annotation See 3d Fortset .p. 35
show subjects concepts


lines 13—12 underline "gelbe ... Blume"
lines 13—12 annotation vars.


lines 10—6 score
lines 19—6 annotation In Henslows List
     considered as varieties: I am nearly sure has been experimented on.

top-marginline 18 annotation Watson in Cybele «seems to» consider them distinct : says perhaps «or probably» 2 species both varying. Refer for experiments t Magazine of Nat Hist V. p.493. & VIII.634 & Phytologist 2.164
line 6 score
lines 10—6 annotation Ask Babington.—


line 15 crossing-out "phoeniceum"
line 13 annotation purpurea
show subjects concepts


line 3 score [`bookmark']


lines 9—7 score
line 8 annotation (a)
bottom-margin annotation (a) Hybrids are affected especially in male organs, with exceptions .


lines 9—1 score
lines 9—4 score
line 9 underline "Liliaceen"
line 9 annotation often mentioned
line 8 underline "und ... Gewächsen"
line 14bottom-margin annotation In these plants, pollen, thogh in appearance good ,yet no impregnation follows ‹[ may be fault of female organs ? C.D]› pollen though swells, does not burst, in water, yet admits it may be owing to female organs , or structure of roots


lines 2—7 score
lines 2—7 annotation (a)
top-margin annotation (a) In this hybrid M. jalapo-longiflora, own pollen more powerful than own concepcion power.


lines 14—16 score
lines 22—24 score
lines 13—28 annotation even in most fruitful hybrids normal number of seeds never produced & always mingled with bad ones.
     compares this fact to result of Hybrid fructifications.


line 19 score [`bookmark']


lines 8—13 score
lines 8—13 annotation Power of fructification in Hybrids always weakened

lines 17—22 score in pale pencil
bottom-margin annotation Speaks of bisexuality of quite exceptional in vegetable Kingdom—


line 17 underline "Dianthus japonicus"
lines 15—19 annotation case of pure species with female organs impotent while male perfect
line 22 underline "Passiflora coerulea"
line 22 annotation so Passiflora
lines 9—8 underline "freien ... Langsdorfii"
lines 9—8 annotation so this

lines 2—1 score
bottom-margin annotation On other side pollen fails, yet female organ quite perfect & potent; in some Dianthus, this happened only with individuals plants.—
show subjects subjects


lines 22—4 score
lines 2—4 annotation The wonderful cases, where in Lobelia, Verbascum & Zephyranthes, pollen wd «not» impregnate own stigma, but wd. impregnate other species; these ‹pol› stigmas being also impregnated by pollen of other species

line 8 crossing-out "9"
line 8 annotation 8/
line 7 crossing-out "10"
line 7 annotation 9/
show subjects concepts


line 1 underline "rustico"
lines 1—5 annotation case in single individual of the Hybrid

lines 20—1 annotation Gaertner has gret advantage that th sexual organs certainly are weakend, as prvding so few seeds

lines 7—3 score
lines 6—5 annotation (B)
bottom-margin annotation (B). Puts this under category, that male less potent than female in each case ; but surely Herberts is more true , viz advantage of crossing. — See to Herbert:—

line 2 underline "Weiblicher ... mit"
line 1 underline "viel häufiger"
lines 2—1 annotation (much more frequent)


line 4 underline "III"
line 4 annotation (B)
top-margin annotation (B) In this III. pollen of Hybrid wd not act on self, but on both parents; & pollen of latter impregnated Hybrid. accounts for this (not as I shd. by advantage of crossing ,& which I still think must hold in Herbert's case), but by believng (& it is probable) that both male & female organs weakened , & cd not act on each other , but only pure parents, or even th Nicotiana on a 3d. species.

lines 4—8 annotation (Is there any parallel III case in pure species? ?)

lines 10—25 annotation These [I, II. & III] cases in Hybrids wonderful parallels to what happens in joining pure species.!!!


lines 1—5 score
line 3 annotation (a)
top-margin annotation (a)    Reurges male organs fail first & most in Hybrids

lines 17—20 score
line 21 score
lines 16—20 annotation says «above» ‹again› analogous with animals
show subjects concepts

lines 13—9 score in pale pencil
lines 13—11 annotation in pale pencil (B)
lines 8—2 score in pale pencil
bottom-margin annotation (B) In dioecious plants not hybrids , in females, male organs sometims imperfectly developed , yet can fertilise ; but in male rudiment of pistil never acquires power of conception.


top-marginline 25 annotation In L. Vespertina, ‹th› in female flower, the rudiments of stamens much smaller than in L. diurna, & consequently only in latter are ‹tr› anthers sometimes found. }
     Does not this well show that a rudiment has something essential & real in it —
     Very Good We can prove Mammae in Male to be a reality.— Wings in insects & Here we can prove in anothr way. Gaertner somewher suggest in Carrot to cut off th fertile flower early & see whether other flowers wd. become fertile . At p 345 «& p. 330» long description of crosses of Dioecious plants
     study it all.— /

lines 14—5 annotation (z)
     Similar changes take place easier in Monooecious than in Dioecious


lines 1—6 score
lines 3—4
top-margin annotation accounts for Hybrid breeding «easier» with parents , than with ‹sef› self by own pollen having less strength; so in cases as below (zz)
lines 8—6 score
line 8 underline "rusticolanceolato-rustica"
line 7 underline "der paniculata"
lines 8—6 annotation (zz)

lines 5—1 score in pale pencil
line 5 at "333" annotation 357
line 5 underline "Lobelia fulgens"
bottom-margin annotation I cannot see how his explanation holds good, for the pollen of L. fulgens (s. 64) did impregnate two othr pure species .


lines 8—13 score
line 13 underline "wie ... hat"
line 13 annotation !
line 10 annotation (a)
top-margin annotation (a) says much experience has shown him that Hybrids, after ‹many› «lines 8—10» generations, have their procreative powers weaker & weaker; & hence cannot be prolonged.— (But then ‹almo› all crossing avoided.—)

line 4 score [`bookmark']


line 2 annotation quite sterile
show subjects concepts

lines 11—15 annotation Some individuals of these Hybrids quite sterile.

line 16bottom-margin annotation Degree of fertility in most Hybrids, except the quite sterile, generally vry variable, even in Hybrids from same capsule & reared alike (This shows how innate, & is opposed [corrected from `composed'] to its being a character of species, as species.)—C.D
show subjects subjects

lines 5—1 score lines 4—3 annotation P
bottom-margin annotation (P.) Hence different accounts by different authors, as follows,—


lines 5—6 score in pale pencil
line 6 annotation A
top-margin annotation A) This unfixedness of fertility of Hybrids, their special character ,& not observed in pure species.—

lines 13—23 score in pale pencil
line 13 annotation (B)
bottom-margin annotation (B) «Some» Hybrids, produce only seed at end , or middle, or (generally) beginning of their flowering, & are at other times sterile.— This a peculiarity, confined (when so pronounced) to Hybrids.—


lines 10—7 (    )
lines 8—7 annotation Does not believe


lines 1—3 score
lines 1—2 annotation (a)
top-margin annotation Cases in pure Dioecious plants of changes in sexual relations

lines 7—12 score
lines 5—9 annotation Female sterility of D. Japonicus transmitted to offspring in Hybrids.

[continues overleaf] lines 7—3 score
bottom-margin annotation Hybrid Plants which produce an extraordinary number of flowers & are quite sterile ; caused by sterility p. 372


line 2 underline "eigenen"
lines 5—10 score in pale pencil
line 7 annotation (a)
top-margin annotation ⸮ In Hybrids crossed with either parent , & thus assuming fertility & th ancestral form , yet fertility variable in «such» individuals; in th successive generations.—


line 13 underline "der ... Liliaceen"
line 13 annotation (a)
top-margin annotation (a) seems to attribute sterility of Liliaceous Plants to state of roots.—
show subjects subjects

lines 5—2 score
lines 5—1 score in dark pencil
lines 5—1 annotation all sterile Cape of Good Hope Oxalis


top-margin annotation My point that plants often sterile & yet not unhealthy not touched on. —
     G. gives only obvious cases of infertility.

lines 12—14 score
lines 12—17 annotation Hybrids in pots more fertile than in open ground.

lines 19—21 score
line 23 score
lines 18—23 annotation In fruitful years more birth from domestic animals

line 9 at "Erschöpfung" annotation exhaustion

lines 4—3 score
bottom-margin annotation More often cause of infertility on male than female side; as in Caryophyllea & Verbascum


lines 13—15 score
lines 16—18 score
line 17 underline "manchen ... unseren"
lines 12—15 annotation cases where pollen good but female organ

lines 20—30 annotation often in Exotics, pollen & female organs are ready at different times, & so can be impregnated artificially ∗
bottom-margin annotation ∗ Would he say that C. Sprengel's facts were due to climatic influences?

lines 10—7 score
lines 10—7 annotation insects less important than wind !!
show subjects subjects

[continues overleaf] lines 4—1 score
lines 4—1 annotation Infertility through long cultivation by layers &c


[continuation] 1 at "Trauben" annotation grape
lines 2—5 annotation disputed by Reichenbach

lines 10—30 annotation Case of wild Verbascums & in pots , with certain flowers sterile & certain fertile; cannot explain. like Kolreuters cases


line 6 underline "männlichen"
lines 6—8 annotation Male Hybrid pheasant sterile

lines 15—19 score
lines 13—20 annotation Morton thinks relation between capacity of Hybridising & domestication

line 5 underline "Fruchtbarkeit"
line 5 annotation (a)
bottom-margin annotation (a) Fertility a fixed attribute of pure species «(in natural conditions C.D)» t01
     Mem. cases of moss not breeding, «(these are probably ‹diociu› Dioecious)»t02 in Hybrids a varying attribute.
t01 - `« (in natural ... C.D)» ' in pale pencil
t02 - `(these are ... Dioecious)' in pale pencil


line 3 at "beschuldigt" annotation charged
line 3 at "Vorurtheil" annotation providence

line 7 underline "chinensi-barbatus"
lines 7—9 annotation fertile according to Kolreuter


lines 12—14 underline "Bei ... Arten"

lines 15—18 score
line 16 underline "immer"
lines 16—18 score
lines 15—20 annotation In pure species artificial impregnation does not alway yield full nmber of seed

lines 11—6 annotation in pale pencil I do not think G. had Green House he always speaks of Ximmer [ie `Zimmer']

[continues overleaf] lines 7—5 score
bottom-margin annotation Hybrids always less seeds than pure parents , as in following examples.—


top-margin annotation See in Beitrage p 598 Lychnis vespertino-diurna gave with own pollen 234 seed .

[continuation] 1 at "40" annotation (24)
line 3 underline "lines 220—280"
line 4 underline "lines 600—800"
lines 1—5 score
lines 1—5 annotation see to Kolreuter about Datura
lines 7—8 score
line 7 underline "192"
line 8 underline "210 ... Samen"
lines 7—11 annotation

lines 7—2 score in pale pencil
lines 6—5 annotation B
bottom-margin annotation (B) Great differences in «different individuals of » same Hybrids & in different years, in fertility striking. —


line 2 score [`bookmark']

lines 14—12 score
line 12 underline "Oenothera ... weinigsten"
lines 14—11 annotation Genera with most fruitful Hybrids


line 1 underline "purpureo-lutea"
top-margin annotation Henschel says this fertile, but reverse quite sterile. See Henslow.
from End Note annotation (p. 387    Digitalis for comparison with Herbert)

lines 13—20 annotation No relation between fertility of pure parent & the facility of uniting, or with these Hybrids having fertility

line 12 at "mithin" annotation consequntly
line 9 at "Beurtheilung" annotation examination


line 1 at "Mass" annotation proportion

lines 5—7 score
lines 5—7 [→
lines 5—10 annotation Hybrids nearly as fertile, but never quite , as pure parents. —

first table annotation The Reverses of these not equally fertile.

line 18 annotation — (K) in Table

line 15 annotation —(ξ G)
show subjects concepts

[continues overleaf] lines 14—9 score
lines 12—10 annotation Some great mistake
lines 14—1 annotation — This not in list
     ←⧟ [to next page]
     not in list !!!
     as repeated at p. 402
     ⧟ (The table is probably wrong)

\7 underline "geringem"
lines 7—1 annotation in little degree fertile, which is commonest case


lines 14—17 score in pale pencil
top-marginline 17 annotation The «abve» numbers show ‹gre› that in Hybrids greater inclination for sterility than for fertility.

lines 9—7 score
lines 14—3 annotation Fertility so variable at different times ,& in different individual, that simple classes of fertile & infertile Hybrids will not do.


lines 5—11 score in pale pencil
lines 5—13 annotation various striking cases of difference fertility , as found by different authors.
line 14 underline "immer total"
line 14 annotation (4) (B)
top-margin annotation (B) Here are 4 cases, in which other authors found fertility whereas Gaertner finds great sterility: was he bad Gardener?
line 8 at "fruchtbar" annotation (1)
line 8 underline "unseren"
line 9 underline "total steril" 11 underline "bei uns"
line 11 annotation (2)
line 11 underline "unfruchtbar"
line 13 underline "bei ... total"

lines 14—19 score
lines 15—19 annotation one year so fertile as to self sow, in next year very sterile.

lines 6—3 score
lines 5—4 annotation P
bottom-margin annotation (P). «Cases of» Hybrids out of same capsule , of different degrees of fertility & some quite sterile. —


line 7 score
lines 8—14 annotation Variability of Fertility cannot be accounted for by luxuriance

lines 17—20 annotation In pure species, when periodically infertile not very luxuriant.

lines 4—3 score
bottom-margin annotation ⧟ (Fertility does not stand in inverse relation to their Luxuriance)


line 10 at "Ableger" annotation in dark pencil layer

line 16 annotation in dark pencil ‹f›
show subjects concepts

lines 4—1 score
line 5 underline "Geum urbano"
line 4 underline "fruchtbar ... unfruchtbar"
lines 9—7 annotation in dark pencil Fertility does not go by genera .


top-margin annotation ☞|| Certainly a priori, one wd. have expected a gradation in fertility of hybrids & old mongrels as Dogs
     In Gaertner Tables there is approach to this.

line 4 score [`bookmark']

line 11 at "nach Maassgabe" annotation in proportion


lines 3—13 annotation Contradicts Wiegman that maternal or paternal types fertile intermdiate sterile.

line 13top-margin annotation Chief conclusions
     (1) Unfixity of fertility in same hybrid. The varying form of th ‹[illegible word]› Hybrid in th abstract which can be divided into following classes.


[continues overleaf] lines 2—1 underline "zusammengesetzten ... unfruchtbar"


[continuation] 1 underline "Dianthus chinensi-caryophyllus X barbatus"
line 1 at "barbatus" annotation male

lines 5—7 score in pale pencil
line 6 annotation in pale pencil (a)
top-margin annotation a| Hence resemblance of Hybrids to either parent no marked influence on fertility.—

[continues overleaf] lines 20—12 score
line 20 annotation (B)
lines 10—3 score
lines 10—9 annotation Examples
line 7 underline "256"
line 5 underline "diesen ... Bastarde"
line 4 underline "absolut unfruchtbar"
line 3 annotation in pale pencil Examples
lines 2—1 score
line 1 underline "49"
bottom-margin annotation in pale pencil (B) These authors think law of relation between fertility of Hybrids & the affinity of parents ; but if we judge of latter by seeds yielded, there is no relation to fertility of Hybrids when reared from them


[continuation] 2 underline "geringem ... fruchtbar"

lines 9—10 score
lines 9—11 score
line 11 double score
lines 8—15 annotation We conclude that fertility of hybrids stands in no near relation to ‹sexual› elective affinity of parents.—

line 25 annotation X
lines 16—19 annotation ⧟ (Yet strong exceptions on next page)

lines 16—2 annotation When plants cross easily both ways, hybrids most commonly fertile. This fertility seems to depend on resemblance in Habitus of parents, but with exceptions


lines 2—12 score in pale pencil
line 6 annotation in pale pencil (a)
line 10 underline "vespertino ... vespertina"
line 10 at "fruchtbarer" annotation yes for see p. 385.
top-margin annotation (a) In Hybrids frm reversed crosses , even when quite like each other , yet fertility not same , & in one case even on one side quite X sterile.
     ☞ Important as shows not in essence of Hybrids.—t01
t01 - `☞ Important ... Hybrids.—' in dark pencil

lines 13—16 score
lines 13—16 annotation Case where reverse cross easy, yet Hybrid sterile

lines 6—2 score in pale pencil
lines 5—4 annotation (B)
bottom-margin annotation (B)    From ‹power› «facility» of union cannot infer fertility of product


lines 9—10 underline "systematische ... begünstige"
lines 8—13 annotation It seems that systematic affinity of Parents favours th fertility of Hybrids . see p. 410.
lines 13—11 score
lines 14—10 annotation Above law it seems has been discussed

lines 7—1 score in pale pencil
line 7bottom-margin annotation see p. 414.    Hybrids from these have remarkalbe fertility & were considered by Kolreuter as varieties. ----
     When we consider these facts we might conclude that fertility of Hybrids indirect relation to affinity of parents


[continuation] 1 underline "Herbert"
lines 5—7 annotation Examples as before

line 12 ]    [ [`bookmark']

table annotation Examples of nearly related species, having hybrids quite sterile

lines 8—4 annotation Most unlike dogs breed & produce fertile offspring.
show subjects subjects

line 1 score
bottom-margin annotation Concludes that likeness «in Habitus» cannot be ground cause of fertility or sterility of Hybrids.


line 12 underline "constitutionellen"
lines 12—15 annotation considers this an unknown element

lines 8—5 score
bottom-margin annotation Repeats that as fertility varies «in Hybrids» from same parents, it belongs to th individual & not to th Kind


line 13 underline "Fruchtbarkeit ... Bastarde"
table annotation considered varieties by Kolreuter from fertility

lines 18—13 annotation (O means not tried by Gaertner)
lines 15—16 at "Datura" annotation ia
lines 17—20 at "Hibiscus; Cheiranthus; Sida; Digitalis" annotation O
line 21 at "Malva" annotation ia
lines 22—23 at "Alcea; Linum" annotation O
lines 7—6 at "Linum; Mirabilis" annotation O

line 15 annotation ?
line 17 annotation —?
line 18 annotation —vars
line 19 annotation — var.?
show subjects concepts

line 13 annotation XX
bottom-margin annotation XX. V. My M.S. p. 19 on Kolreuter , showng that all Botanists agree in thinkng these only vars.

lines 7—6 score
lines 7—6 annotation horizontally crossed not tried
     not tried

lines 9—4 annotation { These 2 are added, though Kolreuter cannot dare to call only vars.
line 4 underline in dark pencil "parum vel"


top-marginline 10 annotation But as several of these are probably vars , it is vry important that G. says not so fertile as pure parents, for we get then a series

lines 3—8 at "Aquilegia; Datura; Lychnis; Lobelia" annotation ia
line 9 at "Matthiola" annotation (k)
line 10 at "Dianthus" annotation ib
line 11 at "Dianthus" annotation ig

line 9 annotation Bentham says var.
show subjects concepts

lines 12—14 multiple score
line 13 underline "ausgezeichneten"
lines 14—15 underline "niemals ... hervorbringen"
lines 3—12 annotation also highly fertile Hybrids
     but yet not so fertile, in any of above cases, as pure parents.

line 12 underline "allen diesen"
lines 13—22 annotation Yet he has not tried all Koelreuters must mean these last alone

line 5 double score in pale pencil [`bookmark']


lines 3—6 score
lines 4—5 annotation (a)
top-margin annotation (a) Agrees with Herberts constitutional doctrine (which I think means only some internal difference.


lines 12—9 score
line 10 annotation a
lines 5—2 score
bottom-margin annotation (a) The most fertile hybrids always lose fertility in successive generations.— Some sterilish plants if artificially fertilised increase in fertility
from End Note annotation p. 418    Ask Author
     There are facts on variation.—


lines 13—19 score in pale pencil
lines 12—30 annotation In 2d & other generations of Hybrids, fertility becomes unstable & often less, so that even parent-pollen will then have little or no effect. This sterility , however, varies much in different individuals & depends especially on the individual

lines 9—7 score
line 8 annotation A
bottom-margin annotation (A) In Mongrels, fertility even greater in second generation, than in first. see further on; for this perhaps implies less fertility in crossing varieties.—

lines 5—2 score
line 1 at "büssen" annotation loose
lines 6—2 annotation Fertility never greater in 2 generation, than in first.


lines 2—4 score
line 3 annotation B
top-margin annotation (B) This decrease of fertility in second generation has been observed in less fertile Hybrids of Nicotiana, & fertile Dianthus hybrids, as in examples given. As this is only second generation cannot be due to want of crossing.—
lines 8—9 score
lines 8—10 annotation So in animals according to Morton

line 20 at "vervollkommnet" annotation perfected
lines 15—26 annotation sometimes fertility increased after repeated artificial impregnation in succeeding generation, but this plant has commonly gone back to either father or mother type

[continues overleaf] lines 12—10 score
line 11 annotation D
bottom-margin annotation (D) Many vry fertile hybrids propagate themselv, with unaltered type like pure species, as in list, but always with decreasing fertility .

line 1 at "barbatus" annotation in pale pencil Kolreuter


lines 2—8 score
line 5 annotation (a)
top-margin annotation (a) The ‹manner› «manner» in which type divides, & goes back, varies much.—

line 13 underline "barbato-chinensis"
line 15 underline "chinensi-barbatus"
lines 13—16 annotation offspring of this varied    greatly &    more than from reverse

lines 15—9 score
lines 15—9 annotation Kolreuter compares these with hybrids crossed with pure parents.


lines 15—21 score
lines 14—20 annotation The exceptional or abnormal hybrids, when fertile, generally produce normal hybrids.

line 8 double score [`bookmark']


line 12 underline "superbo-barbatus"
lines 12—16 score
lines 12—16 annotation with own pollen 4 seeds with pollen of D. barbatus 10. seeds.—
line 16 underline "29 gute"
line 17 underline "67 hervor"
lines 16—17 score
lines 16—23 annotation so again, & thus often.— & likewise so with very fertile hybrids.—

[continues overleaf] lines 8—3 annotation & so with quite sterile hybrids, corolla remains longer when dusted with either parent pollen.—


[continuation] 11—14 annotation examples as last page

line 12bottom-margin annotation When parent & hybrid pollen mixed, latter rendered quite ineffectual, so that no need to castrate; just like when foreign & own pollen applied to a plant, own eliminates quite effect of th foreign.— This Curious.


lines 1—2 annotation (A)
top-margin annotation (A) Pollen of a third kind will sometimes produce more effect, than own hybrid pollen.
line 3 underline "Nicotiana ... gute"
line 5 underline "Langsdorfii"
lines 3—6 annotation 13 seeds with own pollen; langsdorfii 16 seeds.

line 18 at "Umwandelung" annotation transformation


lines 15—19 score
lines 13—30 annotation The pollen of the two parents has no regard to their sexes in th effect thy produce, but that pollen, which has most power of metamorphosis or umwandelung, which will be discussed afterwards, I suppose that pollen which soonest converts hybrid into pure species, produces also most seeds in Hybrid.

lines 18—7 score
line 13 annotation A
bottom-margin annotation (A) Urges again variability of offsprng of self-impregnated hybrids. So mongrels are.—


line 1 at "Aufhebung" annotation abortion
lines 1—2 at "ordnungsmässigen Erzeugung" annotation regular generation
line 4 at "begünstigen" annotation favour

line 12 underline "aufsteigenden"
line 16 at "Anlass" annotation occasion
lines 17—18 underline "väterliche Bastarde"
lines 17—27 annotation Paternal Hybrids
     ----
     are offsprng of pollen of ‹twice used› same species twice

from Front Note 2 annotation in pale pencil p 429    Explanation of Koelreuter “auf-steigenden Grad” absteigended grad p. 451

line 6 underline "2"
line 6 at "atropurpureo" annotation
bottom-margin annotation If this ‹had› hybrid had been crossed with pollen of atro-purpura, it wd have been a “Muterliche Bastard” or “absteigenden” grade p. 451


lines 1—8 annotation by Father pollen more seed generally raised than by Hybrids own pollen, but generally not so many as in first cross of pure parents

lines 15—17 score
line 14 underline "einfache"
line 17 double score
lines 15—17 annotation much unfixedness in this class of Hybrids
show subjects concepts

lines 18—30 annotation Like second generation of simple hybrids, these `Paternal Hybrids” vary much & differ much in fertility, out of same capsule. So vry different results from repeated experiments with same species.


lines 1—3 score
line 9 at "Ergebnisse" annotation results
lines 1—13 annotation The more fruitful hybrids vary less, & go back more to paternal type, but have often reduced fertility, as,
     ----
     examples

line 10 underline "schwächeren"
line 10 annotation (?)
lines 17—7 annotation When thy take less after paternal type & are mch less fruitful, so vary much., — generally under 3 types, in accordance with resemblanc to ancestors & parent.—


top-margin annotation I am not quite sure that these two pages are fully understood
show subjects concepts

lines 8—13 score
lines 8—13 annotation Here the type which came nearest father was most fertile.

line 17 at "unverkennbar" annotation unmistakable

lines 18—25 annotation all sorts of variability in type & fertility

line 2 underline "Stammvater"


line 2 score [`bookmark']

lines 16—20 score
lines 16—25 annotation It is clear that fertility does not always at all increase ‹with› «in» resemblance in succeeding generations, with ‹pa› ancestor

line 12 underline "fünf"
lines 16—12 annotation 5 different types out of this “paternal” hybrid
lines 10—8 score
lines 11—4 annotation Here case of coming near the paternal type with considerable fertility


lines 12—16 score
lines 14—15 annotation A
top-margin annotation A Commonest rule «or appearance» in this stage of conversion ,is that th more th hybrids differ frm mother & approach th paternal type , th more thy suffer in fertility.— Thinks th revrse more probably really th law.

line 18 at "Macht" annotation power

lines 12—6 annotation Female organs recover first their powers.—

[continues overleaf] lines 4—1 score
bottom-margin annotation Fertility in this in «such as»
line 1 underline "Nicotiana rustico-paniculata2"
bottom-margin annotation always vry variable


[continuation] 4—8 annotation Examples of above variability in fertility

lines 10—30 annotation These “paternal” hybrids are when self impregnated, generally more fertile, than in former generation, & of themselves tend to approach the paternal type; ie even when self impregnated & are varialbe in structure.

lines 10—3 score
lines 10—9 underline "in ... Generation"
line 8 annotation B
bottom-margin annotation B    This particular hybrid came by itself more fertile, which he seems to consider usual result of repeated impregnations of own pollen


lines 1—4 score
top-margin annotation Thinks th above like avatismus in Animals

lines 8—9 underline "zweiten Generation"
line 9 underline "zweiten Grades"
lines 9—14 annotation when go back to Mother, not quite & unequally .

line 16 at "entsprechen" annotation agree with
line 10 at "Klage" annotation complaint

line 9 underline "Puvis"
line 9u "Van Mons"
lines 9—4 annotation So these authors wrongly dispute tendency to avatism


lines 8—12 annotation Thinks all variation frm cultivation when free tend to go back.

line 19 at "eintretende" annotation appearing
line 20 at "hinausragende" annotation prominent
lines 19—22 annotation tends more to mother than Father

lines 10—4 score
line 9 annotation A
bottom-margin annotation Happens oftener with some genera than with others ; never in th vry fruitful Hybrids—

line 3 at "Vorschreiten" annotation step forward


line 6 underline "Lavatera ... Generation"
lines 7—11 annotation This first time more to mother in another case more to Father.

lines 14—20 annotation In these going back «‹progeny of› Hybrids» types, Hybrids fertility less, sometim gone, never incresed.

line 13 at "weit gediehen" annotation largely prospered
line 8 at "angebliche" annotation pretended


line 10 at "gesteigerte" annotation enhanced

lines 16—4 score
line 9 at "entschiedene" annotation decided
bottom-margin annotation I believe he here argues that going back of Hybrid offspring, & «of» t01 varieties «not crssd» t02 , is evidence of aboriginal foundation form of species.
     ☞ So it is some evidence —[continues on page 441]— V. p. 455 my Note .Good.
t01 - `« of» ' in dark pencil
t02 - `« not crssd» ' in dark pencil


lines 1—2 score
top-margin annotation Thinks th famous Laburnum case a proof of stability of species & tendncy to go back.

line 9 at "vorherigen" annotation preceding
line 10 at "verwendet" annotation employed

lines 13—3 annotation The occasional approch to father in simple Hybrids or in second generation of Paternal Hybrids, is rarer than th approch to th mother.


lines 8—13 annotation amongst simple Hybrids Those that approach Father are more sterile.

lines 19—20 score
line 19 underline "Lycium"
lines 18—21 annotation These are apt to tend to Father

line 12bottom-margin annotation The Paternal Hybrids in 2d degree which go back to Father have increased fertility. These cases liable to error.


line 6 at "geläugnet" annotation denied
lines 6—9 annotation all the above facts like Avatismus in animals.


line 1 at "Aufschluss" annotation explanation

lines 13—12 apparently unintentional mark

[continues overleaf] lines 3—1 score
line 2 underline "weiteren Generationen"
bottom-margin annotation in pale pencil in very fertile hybrids these goings back to mother or father have not been perceived, so prevented apparently by strength of sexual organs.


[continuation] 1—3 double score in dark pencil
lines 1—3 annotation in dark pencil Law of variation

lines 6—7 annotation A)
top-margin annotation (A) These goig back agree with the Abnormal types, except thes latter are the result of th crossing of pure parents: thy also are very sterile.

lines 17—20 annotation In successive generations more variability

line 13 underline "Dianthus ... carthusianorum"
lines 13—11 annotation These sorts of Hybrids give most varieties.
from Front Note 2 annotation p. 444t01 Definition of Gemischte & zusammengesetze Bastarde & Ausnahmen typus.t02
     p 502    Better definition & examplest03
t01 - `p. 444' in pale pencil
t02 - `Definition of ... typus.' in dark pencil
t03 - `p 502 ... examples' in pale pencil

line 7 underline "väterliche Bastarde"
line 7 annotation D. barbatocarthusian,— carthusian.
lines 7—3 annotation varieties seldomer than «in» last case (‹over› next Page on do)t01
t01 - `(‹over› next ... do)' in dark pencil

line 2 underline "paniculatorustico-glutinosa"
bottom-margin annotation These hybrids always «(with one exception)» approach father (or 2d species) & commonly totally sterile
     Yet I thought thy were sometimes more fertile than with own pollen.


line 16 underline "paniculato-rustica3"
lines 16—12 annotation not to be distinguished from pure N. rustica, but less sterile

line 8 underline "paniculato-rustica4"
lines 8—6 annotation & even in this generation less fertile

bottom-margin annotation Mother ‹Father› pure ‹rustica› paniculata
     Grand ‹Father› Mother
     Gt Gr. Mother (3)
     Grt. Grt. gran Father (4) was paniculata    [drawing]t01
t01 - `[drawing]' in dark pencil
show subjects concepts


lines 1—8 annotation Different species are changed at vry different rates into th paternal type, but this varies in same species


line 3 at "Bewandtniss" annotation condition

lines 8—10 score
lines 3—10 annotation colour of flower does not vary more in later generation than in first, which is different frm other variability

line 15 annotation A
bottom-margin annotation A In som case, especially such as are slow to be converted, th fertility is lessened, especially on male side, even when hybrid has gone back nearly to paternal type.

line 7 underline "rusico-paniculata4"
lines 7—1 annotation Even some of these quite sterile in both sexes .


line 14 underline "Nicotiana rustico-paniculata3"
line 14 annotation A
top-margin annotation A Such Hybrids with own pollen improve in fertility & of themselves go nearer to paternal type.

line 1 double score
lines 1—12 annotation Even some fruitful paternal hybrids in 3d degree were quite sterile on male side.
     Generally with higher degree of Paternal hybridism, so much more fertile.

line 13 at "Bewandtniss" annotation condition
line 12 underline "barbato-japonicus"
bottom-margin annotation In each paternal degree this became more sterile on female side, because it approached D. Japonicus which is naturally sterile on female side.


line 10 at "entspricht" annotation agrees with
from Front Note 2 annotation in pale pencil p 429    Explanation of Koelreuter “auf-steigenden Grad” absteigended grad p. 451


lines 6—10 annotation More fertile than correspndng paternal Hybrids

lines 9—16 annotation varies «more» ‹in some manner› than paternal hybrid x⧟→ chinesi-barbatus ♀ barbatus ♂t01
t01 - `hybrid x⧟→ ... ♂' in dark pencil

lines 15—1 annotation These bore lines 6—15 varieties
     Most of the vars have approached very close to pure maternal type


line 8 double score
lines 10—14 annotation More fertile than corresponding Paternal Hybrids xxx
top-margin annotation xxx. I shd. think caused by female side of Hybrid bing most fertile & this side bing crossed with same type , through a male, caused their greater fertility . & is pretty proof of more fertility on female side.

lines 17—28 annotation But even here amongst those most closely resembling Maternal type, quite sterile individuals are found. Here also male side fails in fertility more than female.

lines 12—9 score
line 11 annotation A
bottom-margin annotation A    In further generations , when self impregnated , become of themselves mor like mother & more fertile.— & less variable


lines 10—20 annotation The more fruitful stick to the hybrid type longer than the less fruitful. Rate of going back varies according to Kind


line 8 underline "zur Stammmutter"
lines 7—18 annotation The law that these (I fancy both paternal & maternal, anyhow the latter) of themselves, self-impregnated go back to type of Mother, most important. (t01 A )t02
top-marginline 5 annotation (A) This makes me believe th reported fact that Mongrels go back: Bell insisted on this to me one day , in regard to Pigeons, & I think Dixon did.
     ☞ So Gaertners remark that this proof of Real Species fails, for applicable to Varieties

lines 1—16 annotation → (But it might be said that one var. was an ursprungliche forme.—)
top-margin annotation in dark pencil Neither var. shd be an aboriginal form.
t01 - `(' in dark pencil
t02 - `)' in dark pencil

lines 8—7 annotation The metamorphosis of one Species into another like a variety into another seems opposed to species being something distinct as Entity.— XX
bottom-margin annotation XX It is argued that th sterility of hybrids , shows that species are a distinct entity, then surely the points in which they agree, may be fairly adduced to show that thy are not essentially different

lines 6—5 annotation Mr Tollets case of Malay Fowls so long affecting breed is case of a var. with strnger tendeny than othrs to go back, like species


line 10 at "Tadel" annotation blame


lines 7—4 multiple score cancelled
lines 6—5 annotation (a)
bottom-margin annotation |||t01 (a) The «rate of» metamorphosis depends chiefly on th species employed; but also in less degree on the variety [This rate has narrow limit.] of the individuals employed; on account of different degree which thy go back in type
t01 - `|||' cancelled


line 16 double score
lines 13—16 annotation The shorter th period of Metamorphosis th less variable

line 19 at "Herstellung" annotation restoration

lines 14—12 score
lines 16—5 annotation In reverse cases, the metamorphosis at different rates, even though the hybrids frm th reverse ‹strng› be alike. Thinks this proof of aboriginal creation.

lines 2—1 score
lines 2—1 annotation A
bottom-margin annotation A    Always approach to type of ancestral form before organs of generation quite perfected; ie these are last restored.


top-margin annotation It is not likely we shd understand th slow restoration of the generative organs, as long as we remain so ignorant regarding th essential action of these organs; & why two sexes necessary.—

lines 8—10 score
lines 11—12 underline "allen ... Ausnahme"
lines 6—16 annotation Not seldom quite like pure parents & yet even quite sterile, specially male organs; sexual organs ‹generally› «universally» in some degree affected

lines 16—12 score
lines 18—12 annotation Law, that male organs not only more easily affected but slower restored.

lines 7—1 score
lines 7—1 annotation do not understand.


lines 17—18 underline "einem ... derselben"
lines 18—19 underline "Nicotiana ... beobachtet"
line 25 underline "Diantus chinensi-barbatus2"
lines 6—37 annotation extreme variability of fertility during th umwandelung, not connected with any law.— Never suddenly appears by a jump. Yet one «almost» exception by Kolreuter, with unusual approch to pure type, and Gaertner one othr case with relatively little approach to pure parent.
     These case show that the gain of fertility ‹show that› is due to peculiarities of the individual.


lines 6—12 score
line 8 annotation A
top-margin annotation (A) As in first generation, decided types (given in last page & p 285) arise close to one parent, so it is evident th number of generations required for metamorphosis must vary much.

table annotation On average


line 10 at "übersteigen" annotation exceed
line 7 at "Ziel" annotation limit
lines 13—7 annotation As far as yet known never requires more than 6 or 7 generations

line 6 double score in pale pencil [`bookmark']


lines 11—19 score
line 11 at "Wahl" annotation choice
lines 12—13 annotation (A)
top-margin annotation (A) Attributes th variability not entirely to th difference of th going back of individuals , but also to variability due to long cultivation, for has not perceived it in th wild-growing , nor in th more fruitful hybrids. —

line 13 underline "ochroleuca ... ochroleuco"
lines 2—1 underline "Der ... umgewandelt"
lines 13—1 annotation In reversed crosses, even when hybrids are alike yet thy are not metamorphosed with equal readiness, which shows some difference in their inner nature ----→ example

bottom-margin annotation ‹Thus Dianthus›
show subjects subjects
show subjects concepts


top-margin annotation Hybrids may be considered as a united brother & sister

line 17 at "nochmalige" annotation repeated


top-marginline 10 annotation Would not “Reduction” good be term for Umwandelung = inversion in Dict. ←
     Absorption into th Father form
     Reduction by the Father or of mother or by paternal pollen or maternal pollen

lines 7—10 score
line 13 underline "Dianthus chinensi-caryophyllus2"
line 16 underline "dritten Grade"
lines 7—17 annotation (B) Fertility of hybrids stands in not01 special relation ‹of› to capacity for metamorphosis:
     Examples,— vry sterile & yet in 3 power almost reduced to D. car.
t01 - underlined in dark pencil

lines 20—24 score
lines 20—24 annotation fertile but require 5 powers.    &c &c

line 2 double score
line 1 annotation other reasons for (B).
show subjects concepts


[continues overleaf] lines 8—5 score
lines 8—5 annotation A
bottom-margin annotation (A) Generally with ‹nearer› «less» “sexual affinity” of first pure parents the Reduction ‹quicker› «slower» , & th reverse with ‹less› more fertility


[continuation] 1—2 score
lines 1—2 annotation A
top-margin annotation Foregong examples show no fixed relation between periods of mtamorphos & sexual affinity of Plants.—

line 8 underline "der ... Typus"
lines 6—15 annotation A species with this power of producng a decided type will reduce a species quicker than in th reverse manner

[continues overleaf] lines 13—10 score
lines 14—9 annotation There is also relation to systematic affinity of species,
bottom-margin annotation Systematic affinity must mean “likeness of characters externally visible.


[continuation] lines 14—6 score
lines 12—9 annotation Examples of last Rule
lines 3—1 annotation Exceptions to


lines 9—13 score
lines 10—12 annotation (Q)
top-margin annotation (Q) The different powers of reduction in hybrids frm reversed crosses, show no fixed relation to “systematic affinity of Parents


lines 11—15 score
lines 12—13 annotation (B)
top-margin annotation (B)    Returns to parent-forms throgh self impregnation are very slow, & require mny generations .

line 23 double score [`bookmark']


lines 6—10 score
lines 6—10 annotation The goings back seldom observed in wild plants when experimented on
line 12 at "428" annotation 3?

lines 13—20 annotation Conclusions
     (1) Facility of Reduction not absolutely depends on sexual or systematic affinity.—
     (2) Returns more often to Mother than Father.

lines 8—6 annotation Not all embryos affected alike.


whole-margin annotation I suppose he wd say there was a hatred in th Vegetable Kingdom to these crosses: perhaps his argument directed against those; like Herbert who believe in hybrid origin of species.
show subjects subjects

line 2 at "hergestellt" annotation restored

lines 16—17 underline "Dianthus superbo-barbatus5"
lines 13—18 annotation In reduced hybrids traces of parental character may be yet discovered.

line 19 at "unläugbaren" annotation undeniable
line 1 at "Beharrlichkeit" annotation perseverance

lines 13—8 score
lines 11—10 underline "unzweideutigen"
lines 12—11 annotation (a)
bottom-margin annotation Thinks th Reduction of species affords “unequivocal” proof that th limits of species confined & fixed. How curious. I can see force in this argument in reductions by self-impregnation.—


lines 1—6 score
line 1 at "Genauigkeit" annotation accuracy
top-marginline 7 annotation Excessive care in prevntng parent pollen, Kept in chamber.—

lines 9—8 annotation The old stories of Grasses changing into each other.—
show subjects subjects

line 4 at "Erwähnung" annotation mention


line 8 at "vorgeblichen" annotation pretended


line 5 at "Vertheidiger" annotation defender
lines 4—6 annotation Hornsuch defender of transmutation.

line 13 at "Haber" annotation Oats
13 at "Roggen" annotation Rye

line 17 [?]u "Berg"


line 18m+score [`bookmark']

line 7 at "käuflichen Linsen, Erbsen" annotation bought lentil Pisum
line 4 at "Kichern" annotation chick-pea
bottom-margin annotation p. 80 Wicken = Vicia sativa Linsen = Ervum lens

bottom-margin annotation Amongst seed of Vetch, some chickpea, which produced 2 vars. like Vetch


lines 18—23 annotation 4 vars of Peas, very slightly different, raised out of bought seed.—
show subjects subjects


lines 8—16 annotation no variously coloured seeds produced & it is clear he wd take pains to prevent crossing
show subjects subjects

line 11 crossing-out "1)"
line 11 annotation 3 / (correct)
show subjects concepts

line 3 at "zusagenden" annotation correspndng
line 2 at "Gedeihen" annotation propsperity


lines 7—11 score
line 8 annotation a
top-margin annotation (a) Remarks that many plants when put out of proper conditions do not vary ,& those that do; their inner nature & power of union suffer much less:

lines 12—16 score
lines 11—16 annotation Cases of change of Form. chiefly in Lecoq
show subjects concepts

line 13, 15—16, 22 crossing-out "[footnote numbers corrected]"
show subjects concepts

lines 16—18 score
lines 16—17 underline "Cruciaten, Liliaceen"
line 17 underline "Salvia"
line 18 underline "in ... behaupten"
lines 16—18 annotation these very fixed

lines 10—8 annotation long cultivated plants as
line 6 underline "Cerealien, Leguminosen"
lines 3—1 score
lines 4—2 underline "Dianthus ... u."
lines 6—4 annotation vary


lines 1—2 score
top-margin annotation same cause makes them ‹vary› easily depart from normal Bastard-type

lines 7—9 annotation varieties tend to go back; no facts given

line 8 at "Einwendungen" annotation objection

line 14 double score [`bookmark']

line 10, 5 crossing-out "[footnote numbers corrected]"
show subjects concepts

lines 12—6 score
bottom-margin annotation Quotes Herbert, that domestic variations do not affect organs of generation


line 13 at "verknüpft" annotation tied together

line 11 annotation There are 6 of these classes.


lines 5—8 annotation Simple Hybrids of one type

lines 10—1 annotation (I cannot think why Reduced Hybrids per patrem are here omitted)
from Front Note 2 annotation p. 444t01 Definition of Gemischte & zusammengesetze Bastarde & Ausnahmen typus.t02
     p 502    Better definition & examplest03
t01 - `p. 444' in pale pencil
t02 - `Definition of ... typus.' in dark pencil
t03 - `p 502 ... examples' in pale pencil

[continues overleaf] 5 score in dark pencil
line 5
bottom-margin annotation (a) & (b) Hybrids alike & so also (c) or Reduced Hybrid per matrem.


[continuation] 9 score
line 11
line 23
line 23 annotation (c)


top-margin annotation (a) It is only th quantity of blood from either side which makes a difference.
lines 4—17 annotation Thus these are alike
     (a)
     But when one factor is more powerful in its influence, then there is a difference, as

lines 12—17 score
lines 12—17 annotation very varialbe & generally vry sterile with exception


line 1 at "Vermischte" annotation mixed

lines 2—3 score
line 2 underline "aus ... Faktoren"
lines 7—8 score
line 19 underline "sind ... steril"
lines 15—19 annotation Excessively variable & generally absolutely sterile

line 11 at "Zusammengesetzte" annotation in dark pencil compounded
line 11 annotation 3 species
     Same as last only mother a hybrid

lines 2—1 score
bottom-margin annotation (So Kolreuter also says)— In type always «(yet a prepotent type in any species has some influence)» go to pure Father: ‹but variable with generally vry little› ,«but» in different degrees.—
     fertility various    generally little. —


lines 4—5 underline "vermittelnde"
lines 3—5 annotation are very distinct frm class 5
show subjects concepts

lines 7—12 score
lines 11—17 annotation In this «sub» class th 3 pure parents are somewhat allied

lines 18—25 annotation very little fertility    in «one» folling case vry considerable fertility
lines 2—1
show subjects concepts


line 7 double score
lines 6—5 annotation (A)
bottom-margin annotation (A) In this second subclass, species are used which will not cross without the intermediate & 3d species, & therefore are very little allied in sexual affinity.— These always most closely resemble pure father. Excessively sterile


lines 13—15 score
lines 12—16 annotation a tendeny to vary even in individual plant

lines 17—24 score
line 20 annotation (a)
bottom-margin annotation This extrem closeness to father vry singular & against ordinay Laws of Hybridism, explained by greater potency of pure pollen of Father , as likewise is shown in th 3d class, in which Mother is pure & yet it seem no leang to either side.


lines 19—11 annotation cancelled It seems that where pollen pure & ovules hybrid, then appr [ie `approaches'?] to pure & less variability
lines 9—4 score
lines 8—6 score in dark pencil
bottom-margin annotation not so variable, apparently owig t th potency of effect of pure parent.—


lines 1—6 score
lines 2—4 annotation (A)
line 5 at "überwiegenden" annotation prepondertng
line 6 double score
top-margin annotation A.    Conclusions (1) that hybrid ovules or pollen cause of variability.
     (2) that th pollen, even of hybrid origin has preponderatng influenc over femal !!! I do not see how this second conclusn


line 4 at "bewährt" annotation deserve

lines 8—16 annotation Here whichever species has the most typical strength, the offspring resembles it — : quite sterile

lines 11—12
lines 10—15 annotation Not one in Table


lines 5—8 annotation 4 species united

lines 5—10 annotation In this the several offspring resembled all four parents

line 12 underline "selten ein Individuum"
lines 8—16 annotation Excessively variable, no two individuals alike. Fertility lost.


lines 1—3 score in pale pencil
top-margin annotation It is clear that the more complicated the unions the more variability ensues.

lines 14—11 annotation Hybrids can be told frm pure only by variability

lines 7—2 score
line 6 underline "schon ... ersten"
lines 5—3 underline "niemals ... Arten"
lines 6—4 annotation repeated over & over again

[continues overleaf] 1 score in reddish-brown crayon


[continuation] 2 score in reddish-brown crayon
line 3 score in reddish-brown crayon
line 8 score in reddish-brown crayon

lines 10—13 underline "Die ... aufgehoben"

line 14 score in reddish-brown crayon

line 15 score in reddish-brown crayon
line 16 score in reddish-brown crayon
lines 17—18 score
lines 17—18 annotation not more subject to malconformation than pure species

lines 18—19 underline "Die ... Hybridität"


lines 14—13 underline "sagt ... seien"
lines 14—13 annotation X
lines 11—9 score
lines 11—10 underline "Sageret ... aus"
from Front Slip annotation p. 521 to p. 524 on germination of Hybrid seed. & all seeds.


line 1 underline cancelled "keimten ... früher"
lines 18—20 score
line 18 underline "11 ... Bastardarten"
lines 18—19 annotation X
lines 1—18 annotation cancelled Period of germination in Hybrid seeds vry various
from Front Slip annotation p. 521 to p. 524 on germination of Hybrid seed. & all seeds.


lines 13—14 double score
lines 13—14 annotation X
line 18 underline "Bastardsamen ... Art"
lines 18—16 double score
line 17 annotation X
lines 13—12 underline "weil ... werden"
lines 11—10 score
lines 11—10 annotation (a)
lines 6—4
bottom-margin annotation The typical strength of a species over th other is shown in affecting period of germination of Hybrid seeds
from Front Slip annotation p. 521 to p. 524 on germination of Hybrid seed. & all seeds.


lines 13—17 double score
from Front Slip annotation p. 521 to p. 524 on germination of Hybrid seed. & all seeds.

lines 16—12 score
lines 16—15 underline "daher ... grosse"
lines 12—10 score
from Front Slip annotation p. 521 to p. 524 on germination of Hybrid seed. & all seeds.


lines 4—1 score in pale pencil
bottom-margin annotation Hybrid seeds do not appear to keep so long as pure seeds


lines 3—8 annotation (3) all vry sterile hybrids are not luxuriant

lines 12—17 annotation (4) These hybrids which are most fruitful are th most luxuriant

lines 14—10 annotation concludes luxuriance a peculiar quality of Hybrids.

line 9 underline "Beschleunigung"
line 9 annotation Hastening


lines 4—10 annotation Hybrids flower earlier ,with exceptions


lines 13—19 annotation innmralbe [ie `innumerable'?] flowers . This is odd. Is it not like double flowers ?

line 11bottom-margin annotation stamens & stigma increase in number sometimes, but not both
     rare exceptions to above, when vry distinct species united/


line 17 underline "diejenigen Bastarde"
lines 13—25 annotation Kolreuter accounts for above by sterility, but doubts as most fruitful ∗ Hybrids, ‹pr› are those which produced most flowers
bottom-margin annotation ∗ Yet these are in some degree sterile


lines 18—5 annotation Because Bees freely frequent quite sterile hybrids, for Honey, thinks no close relation between dissemination of pollen & nectar. ‹If winged insect› Might as well as say elytra not connected with prtectn of wings, becaus presnt in apterus insects
show subjects subjects

line 11 underline "Fruchtungsvermögen"


lines 20—15 score
lines 20—5 annotation Never gives so many seeds as pure parent. /
     As pure species are often sterile sterility cannot be taken as proof of hybridsm


lines 4—15 annotation objects that some quite sterile are only annuals,
     & objects that castrated parents have not life prolonged.

lines 16—18 score
line 17 at "Störung" annotation disturbnc
line 17 annotation (a)
bottom-margin annotation In dioecious plants organs imperfect of one sex. in Hybrids perfect, but functionless

lines 17—12 annotation In crossing hermaphrodite to 2 unisexual plants sexual organs repaired .


lines 18—17 underline "Varietätsbastarden ... Tulpen"
lines 18—17 annotation These vary during life of individuals but then variable flowers

[continues overleaf] line 16bottom-margin annotation ‹This› «In» Hybrid (perhaps only a mongrel iγ) som of the flowers in middle of summer & autumn went back to Mother in flowers


[continuation] 3—5 annotation other cases of above

lines 14—17 score
lines 11—18 annotation Suspect the 2 Tropaeolum only vars, yet vry different.

lines 8—5 score
line 7 at "speciosissima" annotation
\7 at "phyllanthus" annotation
lines 7—6 annotation a
bottom-margin annotation (a)    This hybrid for first three years had angular 5 sided stigma , & then became like Phyllanthus . —


lines 12—14 underline "unverändert ... nigrum"
line 15 underline "Dianthus armeria"
line 19 annotation in brown ink X
line 21 annotation in brown ink X
lines 10—20 annotation cases of Hybrids in which type has kept vry constant , in this case for 10 generations, but with lessened fertility

lines 8—4 annotation Above only examples of progeny of hybrids not varying


line 18 double score in pale pencil [`bookmark']


lines 1—5 score
line 2 annotation (a)
top-margin annotation The tendeny to go back, he argues, wd. prevent new species being formed by variation ; but overlooks my notion of selection picking out th new form adapted to new end.—

lines 4—27 annotation --------→
     Local & constant varieties are different as long as new conditions are present, but change them & the species will go back


lines 1—6 score
line 3 annotation (a)
top-margin annotation (a) as opposed to those who believe genera are made by crossing of species, brings case of Verbascum with species most difficult to distinguish , yet most sterile . — /

lines 9—15 annotation Thinks monstrosities not occuring more in Hybrids than pure species, though Koelreuter did think so. —


lines 1—2 underline "an ... Befruchtungen"
top-margin annotation Has made 1000 artificial impregnations


lines 1—3 score
top-margin annotation Hybrids become double like pure species— Does not seem more apt to be double

line 2 underline "G.F. Jäger"
top-margin annotation Has described double flowers in all classes.


lines 8—12 score
lines 8—12 annotation rare case of double hybrid, if parents single

lines 10—7 score
lines 11—6 annotation sparing & retarded dusting with pollen, most apt to brng double flowers

lines 5—3 score
lines 4—3 annotation (a)
bottom-margin annotation (a)    Hybrids more inclined to double than ‹single› pure species


lines 10—15 score
lines 10—15 annotation It wd appear that this stock was impregnated by Plants 100 yards off —
show subjects subjects


line 7 underline "26"
line 7 annotation 27/
line 14 at "34" annotation 35/
show subjects concepts

lines 13—7 annotation luxuriant growth no doubt necessary for doublng, but some other cause shown to exist

lines 3—1 score
line 3bottom-margin annotation near Hot Spring several Plants double


lines 1—4 annotation cases of wild flower double

line 4 at "angeeignet" annotation appropriated
line 4 underline "eingeprägt" annotation impressed


lines 7—10 annotation It is remarkalbe that vegetative strength owing to sterility does not disturb rest of flower

lines 12—8 score
line 15bottom-margin annotation The Pollen & ovules themselves must have to be modified: the variation is not due to mere mixture of two kinds of cells

lines 3—1 score
line 3bottom-margin annotation vry strange that corolla as altered ‹pe› stamen is not modified ‹p› in Hybrids.

bottom-margin annotation In the second generation of Hybrids we have much variation, which is kind of monstrosity .


lines 7—4 score
lines 6—4 annotation XXX (a)
bottom-margin annotation (a) Some «(Qt01 have thought that single species of genera do not vary (Man !) much, but case of Platanus given .
top-margin annotation XXX    This remark vry curious & bears on what I have shown that large genera var most . I do not know whether remark applied to wild or tame. If ‹latter› «wild as I fancy» all is right. If tame it wd. indicate that my explanation of spreding, & favourable conditions must be superseded by some new law. Could it be tested by Loudon, ascertaing th proportion of genera with single species. by Lindley ???
lines 1—5 annotation «(Shd this rule hold for domestic plants, then we may account for it by variability being necessary to improve plant.)» As I thought of doing with Domstic animals. Wd it be good to take domestic Plants & see proportion of species to genera??? «(or do it all by Loudon. that wd be best)» accordg to Nat. Family & whole Kingdom. /
t01 - `(Q)' in dark pencil

lines 5—10 annotation / Maize has one (or two Molina!?) species) / Rye has only 2 specs / Rice only one ? /

line 12 underline "Tormentilla ... canadensis"
line 12 annotation Hardly vary at all anywhere }—

lines 10—9 underline "Mentha ... pubescens"
lines 10—9 annotation These vary vastly {
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lines 10—4 score
lines 8—7 annotation (B)
line 1 at "Hippocastanum" annotation Aesculus    Horsechesnut
line 1 at "macrostemma" annotation (Red Horsechesnut)
bottom-margin annotation (B) But the Platanus & Pavia have more than one species as far as I can find out


lines 1—9 score
line 9 double score
top-margin annotation Admits th crossng in cultivation must check the Ausartung «of plants» but doubt whether this hold in wild Plants.!!!
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lines 15—18 score
lines 15—18 annotation variation affects every part of Plant.

lines 13—7 annotation crossing of species & varieties an evident cause of variation

lines 4—1 score
line 3bottom-margin annotation variability greater ‹in Mongrels than in Hybrids›t01 owig to mongreling thant02 to external agency
t01 - deleted in dark pencil
t02 - `owig to ... than' in dark pencil


lines 1—3 score
lines 1—3 annotation Van Mons 2 kinds of varieties

lines 14—21 score
lines 14—21 annotation some varieties are constant but crosses of «these» vars. very variable

lines 18—22 score
lines 18—22 annotation White Dahlias not one white seedling

lines 5—1 score
line 3 underline "zum Theil"
line 5bottom-margin annotation all agree that vars. cross & produce partly more fertile ‹Varieties› offspring, than th pure parents.— But exceptions as on next Page


line 3 at "von" annotation ⧟ at p. 87, [corrected from `88'] says these two vars grown in garden always kept pure.

line 3 underline "Cucurbita meldet"
lines 3—4 underline "Zea ... major"
lines 3—8 annotation These unite with great difficulty, but offspring very variable & fruitful

lines 8—10 double score
lines 8—10 annotation in brown ink (Q)
lines 8—10 score
lines 8—13 annotation says some vars. of «(Ask)→» ‹Hor› Dogs, some crosses are more fertile than others /
from End Note annotation (p 577 fertility of dogs.)
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lines 14—18 underline "Nicotiana ... grandiflora"
lines 14—18 annotation K. calls these stabile vars. & Gaertner the followng ; some Botanist consider as species /

line 21 underline "unserer ... fortpflanzen"
lines 20—24 annotation finds like Herbert the vars of Hollyock constant.

lines 15—13 score
line 15 underline "Lychnis"
line 14 underline "phoenicea"
lines 16—13 annotation ‹These species in› (A)
bottom-margin annotation (A) These true species in relation to variability like mongrels, in fertility like Hybrids.— Yet Lychnis is wild & not cultivated Plant see p. 582 at top

lines 11—5 score
lines 11—5 annotation Mongrels like offsprng of simple Hybrids, only more variable, (which surely might be expected C.D)

lines 6—2 score
lines 6—4 annotation case of variability in varieties


lines 1—4 score
line 1 underline "Cucurbitaceen"
lines 2—3 annotation (B)
top-margin annotation In Cucurbitae, on sam plant often two kinds of fruit in shape & flavor
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line 6 at "beurkundet" annotation reported
line 6 at "unausgesetzte" annotation continual

lines 7—8 underline "Gewöhnlich ... vermittelt"
lines 7—8 annotation Important ----- intermediate & this is commonest in close species see p. 283

line 13 underline "früher"
line 16 double score
lines 9—20 annotation The uninjured & often increased fertility of mongrels, accounted for by luxuriance of Hybrids (I do not see this) & says he find garden Plants varied from crossing vars. are earlier than ordinary vegetables. (The earliness is hardly same as greater fertility.) () (ask)

line 11, 10, 9, 8, 7, 6, 5, 2, 1 ✔
11— annotation on account of greater «but not equal to pure species» fertility Kolreuter considered these as only vars: ‹vars.› — G. thinks «(from actual experiments)» «‹frm exprmnts›» only th Hyosciamus as true varieties; because by cultivation one turned into other
     (Steudel makes agrestis a Synony of H. niger G. says thy turned into each other)


lines 4—6 score
lines 6—8 score
lines 5—6 underline "Verbascum ... luteum"
line 7 underline "Bastardverbindung"
lines 4—8 annotation same conclusions regarding these /

lines 9—11 score
line 10 underline "Bastarde"
line 11 underline "absolut"
lines 8—14 annotation says fruitfulness of Hybrids not absolutely ‹connected with› proof of parents bing only varieties
lines 9—14 annotation ask author, I fancy means only some fertility.

line 1 score
line 1 annotation X
line 2bottom-margin annotation Examples of Hybrids vry fertile but not as fertile as pure parents
     These are not hybrids, but merely th union of two pure species .t01
t01 - `These are ... .' vertically crossed


line 6 annotation The Hybrid frm this quite sterile

lines 20—10 score in pale pencil
line 20bottom-margin annotation seems to admit quite the crossing of varieties left to themself, & may be cause of return of vars to parent forms. /
     says mongrels in their variation in successive generations may be classed like the Hybrids, which he has classed.


lines 1—4 score
line 2 annotation (a)
top-margin annotation Says «besides infertility» mongrels differ frm Hybrids in ‹di› varying in th first generation whereas Hybrids vary in 2d or in paternal & maternal reductions. He gives no case of wild varieties ,when first crossed varying more than mongrels; at least I cannot remark any.
from Slip Attached 1, Side 1 annotation in pale pencil Does he give any case of two wild varieties when crossed , producing a more variable offsprng than ‹pur› true species ? in first generation , because this difference in variability he makes an important distinction in vars & species (p 581)

line 6 underline "andere ... Bastarde"
line 9 double score
lines 5—9 annotation in dark pencil in other respects like Hybrids only more like to pure species.

lines 10—25 annotation He evidently considers these vry important characteristic differences (just repeating p. 273) between ‹Hybrids & mongrel› crossing species & varieties.
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[continues overleaf] lines 3—1 score in dark pencil


[continuation] 1—4 score
lines 1—4 annotation merely repeats p. 577

lines 9—10 annotation more accessilbe to impregnation of othr vars
lines 12—13 annotation More tendeny to revert to parent form
lines 14—17 score
lines 14—17 annotation more variable
line 18 underline "gewöhnlich"
lines 18—19 annotation commonly more fertile

lines 7—1 score in pale pencil
line 4 annotation a
bottom-margin annotation (a) Lecoq states great variability in Iris, supported by observations of Berg, hence suspectst01 tht there may be variety-bastards — So necessary to show no need of crossng look at Potatoes & Maize & Rice !!! ‹Horse›
t01 - underlined in dark pencil


line 15 score [`bookmark']


lines 4—7 annotation The smaller proportion of Hybrids are “intermediate”


line 2 at "Wimmer" annotation ∗ 15/    all follwg numbers wrong
line 10 crossing-out "25" annotation in dark pencil 26 /
bottom-margin annotation see Corrigenda


line 16 crossing-out "53"
line 16 annotation 54 /
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line 2 underline "N. rustica und"
line 2 annotation Natural Hybrids
lines 5—8 score
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lines 15—11 score
line 13 annotation (a)
bottom-margin annotation says not knwn how lng & in what limits keep true.
     but wheat shows how long can be preserved under same conditions .
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line 4 underline "paniculato"
lines 3—5 score
lines 3—6 annotation He puts Mother first & Father after, some have followed an opposite course.
from Front Note 2 annotation p 602 Mothers name first
from Front Note 2 annotation in pale pencil p. 602 Nicotiana «‹Female›» glutinosa ♀ «Mother» — «male ♂» perenne «Father»

line 5 double score [`bookmark']


lines 8—3 score
bottom-margin annotation Thinks th facts of Hybridisation shows that original species forever remain true


line 7 at "XXXVII" annotation in brown ink all (Q)
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lines 4—1 score
bottom-margin annotation as varieties can generally be propagated, as known for centuries, any alterations, if they ever occur, require careful observations.—


[continuation] 6—10 annotation sometimes less fruitful, sometimes more.— /
line 16 underline "vollkommenere"
line 17 underline "zahlreichere"
line 18 underline "Geschmack ... Früchte"
lines 16—22 annotation seedlings generally bear more perfect & more numerous seeds than when grafted.
lines 13—7 annotation sometimes life rendered longer, sometimes shorter lines 4—1 annotation longer in foreign trees

line 10 at "3" annotation —(?)
line 3c "4" /@line 33 annotation 5
line 1 double score
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lines 10—13 score
lines 10—15 annotation evergreen oak grafted on common cast leaves & Daphne laureola flowered in winter

line 23 at "Mandel" annotation almond
line 23 at "Pflaumen" annotation plum
lines 15—22 annotation effect of one Pear grafted on an earlier kind was to make it actually later!

line 10 at "10" annotation 11 /
line 10 annotation ‹(10 [illegible word] is right)›
line 4 at "12" annotation 13
line 3 at "13" annotation 14
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line 10 at "Mispel" annotation medlar
line 6 at "Pfirsichblüthen" annotation peaches


line 11 underline "Oleander"
lines 14—10 annotation cases of mottled leaves affecting th Stock.


lines 5—9 annotation Even th wood keeps distnct at place of grafting.


[continues overleaf] 2 underline "allein ... vermischte"
line 5 underline "selbst ... zu"
lines 1—9 score in pale pencil
lines 1—14 annotation a statement that two kinds of grapes — branches split & joined longitudinaly produced striped fruit & crossed foliage . G. does not believe.
lines 8—5 annotation other similar cases

line 11 crossing-out "45"
line 11 annotation 46 /
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line 5 underline "53"
lines 5—25 annotation Case of sport in common Laburnum with flowers like C. Adami
     Is not this like the orchid case? Were they sterile? The sport & parent in Austrian Bramble are sterile.
     (Herbert has shown are sterile, in Hort. Jounal)t01
t01 - `(Herbert has ... Jounal)' in dark pencil

line 18 at "Trauben" annotation grapes

[continues overleaf] b annotation (B) He is dreadfully puzzled about th Laburnum case & says not analogous to anything known


[continuation] 4—16 score in pale pencil
line 12 annotation B

line 7 at "Mandel" annotation almond
line 7 at "Pfirsichbaum" annotation Peach
line 6 at "Pfirsichs" annotation Peach
line 6 at "Pflaumenbaum" annotation Plum
line 6 at "Birnen" annotation Pear
lines 14—4 annotation Power of ‹budding› «grafting» much larger than of hybridising; even vry different genera
     (A) (t01 It makes it the more remarkable that certain vars. shd. not do well together. )t02

bottom-margin annotation (A) I think I have heard it said same Family.    Syringa Fraxinus Olea Chionanthus } all Oleaceae
t01 - `(' in dark pencil
t02 - `)' in dark pencil

line 11 crossing-out "55" @11 annotation 6 /
line 9 crossing-out "58" @9 annotation 9/
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lines 2—8 score
top-margin annotation The relation of th different kinds which can be grafted on same stock is vry different from the relationship on which hybridisation depends

line 8 double score [`bookmark']


line 11 crossing-out "60"
line 11 crossing-out "61"
line 11 annotation 1 / 2 /
lines 7—12 annotation A certain affinity necessary beyond doubt.

line 16 at "Dichtheit, Schwere, Biegsamkeit" annotation compactness Weight flexibility
line 15 underline "organischen Structur"

lines 9—8 underline "die ... Individuen"
lines 11—5 annotation The above influences not only possibility of graft, but ‹fertility› fruitfulness & duration of life

lines 5—1 score in dark pencil
line 6 underline "Familien-Affinität"
bottom-margin annotation ⧟ Family affinity, though greater difference between the graft & stock in wood, yet permits th graft.


lines 8—10 score
line 8 at "Birnen" annotation Pears
line 9 crossing-out "63"
line 9 annotation 4
line 10 crossing-out "64"
line 10 annotation 5
lines 8—9 underline "schlagen ... Diel"
lines 8—9 annotation in dark pencil (Q)
lines 3—13 annotation great difference in powers of grafting. Pear & Apple though allied will with difficulty graft.— Difference in reverse case

line 18 annotation Will not hybridise.
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lines 15—12 score in pale pencil
lines 15—12 annotation can be grafted but not hybridised


line 11 crossing-out "70"
line 11 annotation 71 /
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bottom-margin apparently unintentional mark


line 4 double score [`bookmark']


line 15 score
line 15 underline "Herbert"
lines 9—15 annotation Merely how thy worked
     (p. 354. 369. 374)
     See —


lines 8—7 score
lines 8—7 underline "Unkenntnis ... Gewächsen"
bottom-margin annotation Ignorance of process of fructification in some plants has caused ‹m› failures


lines 9—13 score
lines 9—13 annotation has never seen ill effects from castration, except when all castrated.

lines 9—5 score
line 7 annotation (a)
bottom-margin annotation Dichogamous plants less ‹liabl› capable of hybridising ; & very liable to crypto-hermaphrditism .—


lines 1—20 annotation in dark pencil (Can the pollen of another individual or var overpower own pollen?)—

lines 8—9 score
lines 8—12 annotation Best generally to castrate at moment of opening of flower.

line 19 underline "Leguminosen"
line 20 underline "Malvaceen"
lines 18—22 annotation Necessary have cut or open or partly or wholly cut away petals

lines 16—14 score
lines 15—14 underline "Leguminosen ... Campanulaceen"
lines 16—12 annotation (a) (Quoted)t01
bottom-margin annotation (a)    Anther ‹ready› ripe before opening of flowers in these Families & the fructification takes place not only some hours, but even days before flowers open.    Then how do Cruciferae «& Peas» cross ??
t01 - `(Quoted)' in dark pencil

line 15 at "Onagrarien" annotation (Oenothera Epilobium Fuchsia Clarkia)
lines 9—7 annotation (Lecoq says pollen of Fuchsia not shed for 3 days after flower opened)

lines 5—1 annotation / even whole corolla can be removed without injury to seeds /


line 16 annotation Pincers


lines 1—2 score
lines 1—4 annotation Pistil grows in Geum after impregnation.
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lines 15—11 annotation By mny plants pollen & ovarium not ready at same time

lines 7—4 score
lines 7—4 annotation Impregnate early, because Pollen keeps its strength

bottom-margin annotation Stigma generally ready when flowers open, but sometimes not ready for some time afterwards


lines 2—10 score
line 6 annotation (a)
top-margin annotation (a) The impregnation with own pollen, ‹fructi› fertility always greater than in any Hybrid, & equal or at least near natural fertility, but sometimes less.— Really this accounts for th (i a) of Hybrids.—

line 10 double score [`bookmark']


line 1 crossing-out "56"
line 1 annotation 57


line 1 at "Isoliren" annotation ⧟ Isolation only superfluous in exotic plants where only one presnt.—
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lines 4—5 underline "Die ... Nothwendigkeit"
lines 2—8 score
lines 2—12 annotation speaks of th absolute necessity of isolation (& so does Lecoq) which all shows how muc crossing goes on.

line 8 double score [`bookmark']

lines 14—16 annotation Cutting off all flowers injurious


line 14 crossing-out "65"
line 14 annotation 6
line 18 crossing-out "66"
line 18 annotation 7
line 19 crossing-out "67"
line 19 annotation 8
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line 3 double score [`bookmark']


line 18 crossing-out "68"
line 18 annotation 9
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line 5
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line 4 underline "ganzen Habitus"
line 5 underline "M. longiflora"
line 14 underline "5,2"
line 16 underline "12,5"
line 17 underline "3"
line 15 annotation intrmdiat

lines 11—5 score
lines 10—9 underline in dark pencil "Farbe ... Jalapa"
line 8 underline in dark pencil "Grösse ... kamen"
lines 8—1 annotation seed of this Hybrid returned to two distinct parent forms.

bottom-margin annotation in dark pencilvertically crossed & so in Maize I am nearly sure 2. vars «of seeds» in Mongrels.


lines 18—15 annotation fertility varies much in differnt experimnts.


top-margin annotation in dark pencil (Get Hooker to read over this list)

top-margin annotation There are important facts ‹not› in this Table not noticed in my abstract or results.

lines 3—7 annotation in dark pencil instances of series of fertility

lines 12—13 annotation Count how mny pure species have (K) when self impregnated

lines 12—13 annotation in dark pencil See whether any difference in two vars., I have seen to Verbascum
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lines 21—23 annotation Mothers name first

line 12 annotation succeeded with by Kolreuter

line 11 at "coerulea" annotation = arvensis Loudon Cat
line 10 at "phoenicea" annotation = arvensis Steudel
lines 15—3 annotation Herbert succeeded, see p. 653 «I do not think same species» /
     are these th English specis?
     / (yes.) }

line 8 cols 3 & 4 underline "9 | 9"

line 1 col 3 annotation (no of flowers)
line 1 col 4 annotation (no of fruit)

line 2bottom-margin annotation (See how many genera not result; & genera, I believe with close species : this bears on vars.)
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line 26bottom-margin annotation I shd. trust this more if (see case p. 706) more information given ⸮ of result of self impregnation
     The vry near to approch to (K) & yet th rarity of actual (K) makes me think th effect of artificial fecundation .
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top-margin annotation in dark pencil Hooker thinks tht probaly Canadensis & atropurpurea, are merly synonyms «no, see p. 469 middle»t01 : Hooker thinks Canadensis & vulgaris distnct
t01 - `no, see ... middle' in pencil

line 5 at "atropurpurea" annotation Siberia
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lines 8—9 annotation var. Hooker — (of vulgaris) (⸮⸮) —
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line 9 at "viridiflora" annotation (Steudel ‹?› var of atro-purpurea)
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line 14 underline "2" in dark pencil
lines 10—17 annotation in dark pencil ⧟ This means frst hybrid crossed by Father

line 22 annotation (What Ask author)t01    Steudel makes var of atropurpureat02
t01 - `(What Ask author)' in dark pencil
t02 - `Steudel makes ... atropurpurea' in pale pencil

lines 26—27 annotation — var of vulgaris —
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line 28 annotation var of atropurpur ⸮ —
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line 13 annotation cancelled ⸮—
line 10 annotation cancelled ⸮—

line 3 annotation var. of vulgaris
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bottom-margin annotation viridiflora is a Siberian species of Pallas
     var. of atropurpurea according to Steudelt01
t01 - `var. of ... Steudel' in pale pencil


line 3 score
line 1 annotation var of vulgaris
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lines 12—15 score
lines 12—15 annotation Steudel makes synonyms
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line 13 annotation Prop. Poll
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line 22 annotation in pale pencil — Prop pol
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line 19 unmarked
top-margin annotation ⧟ In this country Hooker says, C. littoralis or maritimus is considered a vy close, but distinct species

line 19 annotation wild var
line 17 annotation wild var.

lines 23—3 annotation Dr Hooker considers undoubtedly vars. except Italicus /
     In text p. 197 & elswhr C maritimus is spoken of as crossing with C. Behen
     must be a misprint. anyhow C. Maritimus = S. inflata according to Steudel.—

line 15 at "italicus" annotation (Vry different Hooker)

line 18 annotation in pale pencil —Prop Poll

line 17 score in dark pencil
line 17 cols 3 & 4 underline in pale pencil "5 | 5"
line 14 score in dark pencil
line 14 cols 3 & 4 underline "9 | 8"

line 14, 12, 11, 10, 4 score in dark pencil "i α"
line 3, 2 score in dark pencil "a"
line 1 score in dark pencil "i β"
lines 15—1 annotation All these i α &c are frm crossing varieties.
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bottom-margin annotation Cucubalus alpinus ― behen angustifolius ― behen latifolius ― littoralis «or maritimus»t01 } = Silene inflata, Steudel    Some authors think Silene italica, ― pilosa
     There is nothng in London Catalogue to make me doubt this conclsn
t01 - `or maritimus' in dark pencil
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line 19
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line 25 annotation = incornis of Kolreuter
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top-marginline 24 annotation Asa Gray considers the D. tatula as var of D. Stramonium & introduced into America    Dr. Bromfield in Phytologst says he has tried evry gradation betwen these two forms, & yet here not fertile (K). This then is case of some sterility, if we are to trust th same class of facts as we infer sterility from.—

lines 22—29 annotation See p. 385 for degree of sterility of D. tatula & stramon

line 23 annotation Art.
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line 13 annotation = plumaria Linn.
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line 21 annotation in dark pencil — Prop . Poll


line 17 annotation in dark pencil — Prop. Poll.


line 18 annotation Croatia
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line 11 annotation ‹nati›
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line 1 annotation perhaps var.
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top-margin annotation [ p 225. Much important on reciprocal crosses in Digitalis. ]
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line 11 annotation var of last Lindly make
line 12 annotation ‹probably› «perhaps» vars.
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lines 17—18 annotation some think var of 2 last
lines 16—26 annotation
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line 17 annotation ambigua of Kolreuter p. 175. ‹m› ambigua anyhow probably distinct
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line 19 annotation — ‹probaly› «perhap var» var
line 20 annotation — ‹prob› perhp var.
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line 19 annotation var. of ferruginea acc to Lindly
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line 2 at "spicatum" annotation = angustifolium Steudel
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line 7 col 3 underline "6"
line 7 annotation — Prop pol
line 8 cols 3 & 5 underline "2 || a"
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lines 7—16 score
lines 6—12 annotation crossest01 more fertile than with own pollen.
line 11 annotation —yet Newman say quit fertile
line 16 cols 3 & 4 underline "3 | 3"
line 16 annotation — Prop polln.
t01 - `crosses' in dark pencil

line 22 annotation — Prop. Poll.

lines 8—4 score deleted in dark pencil
line 8 annotation ----t01 Dr Salter Bell says quite fertile Phytologist
t01 - `----' in dark pencil
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line 18 score in dark pencil

line 19 annotation = niger
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line 19 at "pallidus" annotation = niger
line 18 at "agrestis" annotation = niger
line 18 annotation — Probaly Vars.

line 13 at "pictus" annotation = niger ?
line 12 at "pallidus" annotation = niger
lines 12—11 annotation — perhaps vars.

bottom-margin annotation p. 578. G. says agrestis = albus as knwn by experimnt & Steudel makes albus distinct —


line 10 annotation = undulata
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lines 20—19 score
line 20 annotation ‹I think› { Herbert p. 345 succeeded & they sowed themselves.
line 19 cols 3 & 4 underline "22 | 10"
line 19 annotation — !

lines 13—12 score
line 12 score

lines 8—7 score
lines 8—7 annotation Prop. Poll


line 13 annotation cancelled —X

line 7 annotation —X Prop poll
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lines 7—8 annotation cancelled — Pr

line 11 score
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line 18 annotation Prop. pollen

line 13 annotation Prop Pol

line 10 score
lines 10—2 annotation This is speciosa fertile according to Herbert p. 346


line 14 annotation = sylvestris
lines 15—20 score
lines 15—21 annotation see my slip of Paper about Synonyms
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line 15 annotation — Prop polle
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line 18 annotation var. self im

line 20 annotation = dioica
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line 23 score
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line 24 annotation = Silene viscosa
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line 28 score
line 18 score
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line 13 annotation = Silene
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bottom-margin annotation It is evident from Steudel that Silene, Lychnis & Cucubalus all most closely allied


line 9 annotation — p 385. contradicted

lines 16—12 double score
lines 16—9 annotation Here it is evident that first cross normal
lines 16—13 annotation — Prop poll

lines 12—5 score
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line 4 cols 3 & 4 underline "12 | 8"
lines 1—7 score in pale pencil
lines 2—5 annotation (What differences)

line 19 cols 2—5 underline "1839 | 1 | | a"
line 20 annotation —!!
line 24 score
line 24 cols 3 & 4 underline "12 | 4"
line 25 score
line 27 score
line 30 score
line 23 score in reddish-brown crayon
line 30 score in reddish-brown crayon
lines 18—32 annotation (This very important) see Koelreuter about this.t01
t01 - `see Koelreuter ... this.' in dark pencil

line 18 score
line 19 annotation X    Loudon ten week stock O
line 16 annotation X    smooth —|
line 15 score

line 7 score
line 7 annotation Prop. poll.

line 1bottom-margin annotation These seem distinct ‹but have been sometime confounded›
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line 2 score
line 2 annotation Kolreuter raised them

line 4 annotation —?
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line 25 score
line 25 annotation Sageret raised thm p 35

line 5 score
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bottom-margin annotation according to Steudel «nearly» all these are true species of Nicotiana
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line 26 score
line 26 annotation f
line 29 score
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line 15 score
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line 9 annotation — Prop P.t01
t01 - `Prop P.' in brown ink
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line 4 score
line 9 score
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line 27 annotation = suaveolens Lamarckii acaulis
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lines 22—23 score
lines 22—23 annotation p. 168 some authors think vars.

line 19 col 3 underline "10"
line 18 col 3 underline "6"
lines 19—18 annotation — Prop

lines 20—9 score in pale pencil
line 12 col 3 underline "4"
lines 12—10 score
lines 12—10 annotation others have succeeded lines 18—13 annotation More fertile than with own pollen /


line 6 at "Pen" annotation t
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line 9 annotation in dark pencil or coccinus
line 16 annotation coccinus
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line 20 annotation = vidacea St
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line 22 annotation see Herbert
     p. 379 . More fertile than either parent


line 1 col 3 annotation in dark pencil — nmber of Flower

line 1 col 4 annotation in dark pencil — of Fruit

line 5 annotation in pale pencil Prop. Poll

line 15 annotation in pale pencil Prop pollen

line 22 annotation in pale pencil vars fertile

line 25 annotation primrose
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line 24 annotation (Florist var of Oxlip.)—
line 24 annotation Not normal or K
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line 23 annotation Oxlipt01    elatiort02
lines 23—21 score in dark pencil
t01 - `Oxlip' in dark pencil
t02 - `elatior' in pale pencil

line 20 annotation in dark pencil Cowslip —
lines 20—19 score in blue crayon

17—15 cols 3—5 crossing-out "4 ... a"

lines 16—3 annotation p ‹721›.
     p. 247 it is evident that he did cross elatior & officinalis. Table not correct.

line 14 annotation Oxlip —
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lines 13—12 {

11—10 cols 3—5 crossing-out "4 ... a"

lines 9—8 {

lines 7—5 {
line 7 annotation cowslip—
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lines 4—1 score in brown ink
line 4 annotation cowslip    Elatiort01
bottom-margin annotation ⧟ If this be elatior calycantha, most stranget02
t01 - `Elatior' in dark pencil
t02 - `If this ... strange' cancelled


top-margin annotation I see he has not tried Primula proprio polline

3—4 cols 3—5 crossing-out "5 ... a"

line 5 annotation Cowslip
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line 6 annotation primrs
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lines 6—7 score in blue crayon

8—9 col 3 crossing-out "13 ... 11"

line 10 annotation Oxlip var

line 11 annotation Oxlip.
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line 16 annotation = floridum
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line 16 annotation —Prop. poll.
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line 15 annotation Silene inflata —
line 14 annotation S ----
line 13 annotation S. pilosa
line 12 annotation S. italicus
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line 26 annotation Prop Pollen

line 10 annotation Prop. pollen
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lines 9—1 score in pale pencil
bottom-margin annotation I see Moerch considers same species p 549 Gaertner says perhas only varieties


top-margin annotation It is impossible to make out whether vars. albus & luteus are put first & second on principle or by chance

line 3 score in reddish-brown crayon
lines 3—4 annotation ‹make out colour of real species.›    Yellow ‹?› yes see p. 180

lines 5—7 score cancelled

line 10 annotation ‹— thapsus ?›
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lines 8—14 annotation vertically crossed Colour ? If Yellow half agrees & opposed to rule of vars. of same colour most opposed

line 20, 26 at "albo" annotation agrees with
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lines 30—34 annotation 1845 [-] 1827 [=] 18

line 14 at "flore luteo" annotation Blattaria
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lines 8—4 score
line 8 annotation Colour ?
bottom-margin annotation Steudel make = virgatum, which is yellow

line 1 col 3 annotation Xt01 [total] 286
line 1 col 4 annotation [total] 161
t01 - `X' in reddish-brown crayon
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lines 3—4 score
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line 9 annotation yellow
line 11 → cancelled
line 13 → cancelled
lines 11—13 annotation opposed to rule

line 17 annotation Probably yellow, both parents being yellow
line 20 score

lines 17—37 annotation in dark pencil Even Babington admits there are 2 coloured vars of V. lychnitis

line 29 score
line 28 score
line 25 score

line 24 crossing-out "luteo ... H"
line 22 crossing-out "Thapsus ... H"
lines 22—19 annotation These lines merely guiding
line 17 crossing-out "speciosum ... a"
line 7 crossing-out "― cuspidatum ... h"
line 4 crossing-out "Phlomoides ... H"
line 3 crossing-out "thapsiforminigrum ... α"
line 2 crossing-out "elongatomacranthum ... α"

line 20 col 5 crossing-out "f α"
line 15 col 5 crossing-out "f α"
line 13 cols 5 & 6 crossing-out "k | H."

line 10
line 8
line 6
line 5
line 1

line 7 at "cuspidatum" annotation yellow

line 1 col 3 annotation [total] 226 Xt01 ‹182›
line 1 col 4 annotation [total] ‹119› 142
t01 - `X' in reddish-brown crayon
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line 1 score
line 1 annotation Colour see Index.
     not in index

line 19 score

line 25 annotation yellow
line 28 score cancelled
line 28 annotation [illegible word]

line 21 annotation yellow
line 15
line 10

line 10 crossing-out "luteo ... 2"
line 10 annotation ?
line 9 crossing-out "albo ... 3"

line 1 col 3 annotation Xt01 [total] 234
line 1 col 4 annotation [total] ‹72› 83
t01 - `X' in reddish-brown crayon
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line 3 annotation yellow
line 5
line 12
lines 7—8 annotation opposed to rule

line 11 crossing-out "albo ... β
lines 11—12 annotation why luteo put first?
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line 15 annotation Purple

line 24 annotation yellow

line 20 crossing-out "1828 ... f"
line 19 crossing-out "1828 ... 3"
lines 20—17 annotation why luteo put first
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line 1 col 3 annotation Xt01 [total] ‹179› 201
line 1 col 4 annotation [total] 68
t01 - `X' in reddish-brown crayon
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line 1 col 2 annotation in dark pencil — years
line 1 col 3 annotation in dark pencil — no of flwrs
line 1 col 4 annotation in dark pencil — nmber of seed
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line 1 annotation Light yellow
line 2
line 9

line 19 annotation Yellow
line 23
line 28

line 13 col 3 annotation Xt01 [total] 138
line 13 col 4 annotation [total] 84
t01 - `X' in reddish-brown crayon
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line 11 at "numulariaefolia" annotation = serpyllifolia
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lines 8—1 score
lines 8—1 score
lines 8—1 score in brown ink
lines 8—1 annotation vars yet all i β. & i γ.
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line 1 col 3 annotation [total] 22
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line 9 score
lines 4—11 annotation at Hort Soc ?
     account of experiment with Peas, see to this. }—

from End Note annotation Books of great importance to Refer to
     Note 62 — 67 — 17 } p. 734 of this Book—

line 15 score
lines 13—18 annotation Berg Read Variation of Leguminosae.
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lines 2—1 annotation in dark pencil Read
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lines 28—29 double score
line 27 annotation wrng


line 11 score
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line 20 annotation ‹138—›
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line 17 score
lines 18—14 annotation See to this for Blyth
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line 5 annotation [illegible word]
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lines 11—13 annotation / (Read) /
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lines 16—18 annotation Herbert {
lines 16—18 score
lines 16—18 annotation ‹Herbert›
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line 19 annotation — Worth reading or consulting
     p. 145 of Book


lines 3—9 score
lines 1—9 annotation seeds retaining long vegetating power

lines 10—11 score
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line 18 score
lines 17—21 annotation Omalius disputes vars going back

line 15 score in reddish-brown crayon
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line 22 score
line 22 annotation /See to this
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lines 5—4 score
line 3 score
line 3 annotation in reddish-brown crayon X
bottom-margin annotation || Read Girou. on vars. of Cucurb crossng
     Sageret do not crossing


line 11 score
lines 13—11 annotation X Read
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lines 13—14 score
lines 13—14 annotation X
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line 16 annotation X
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line 3 score
lines 1—5 annotation on curious sport in Oenothera    /Read

lines 8—7 score
lines 12—5 annotation Read — on th 2. Anagallis being vars.
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line 10 score
lines 14—15 score
lines 10—15 annotation Read Beitrage    Read
line 3 annotation /Read
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line 14 annotation horizontally crossed Read
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line 2 score
lines 1—5 annotation Zuccarini on sterility of Oxalis from C. of Good Hope
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line 10 score
lines 9—10 annotation
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lines 13—14 score
lines 13—14 annotation Read
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line 16 score
line 16 annotation Read
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line 18 score
lines 17—18 annotation Read
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line 27 score
line 27 annotation Read
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line 30 score
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lines 10—9 [
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lines 6—8 { cancelled
line 8 annotation cancelled ? line 54 score
line 54 annotation
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lines 9—12 score
lines 6—12 annotation cases of transformd plants p. 500 text
lines 9—10 double score in dark pencil
line 12 annotation in dark pencil nothing

lines 13—14 score

line 1
top-margin annotation Perhaps worth reading «But opposed to much alteration»


line 11 score
lines 12—11 annotation p 540 ✔

line 18 annotation X
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line 19 score
line 19 annotation read
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line 12 score cancelled
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line 11 score
line 11 annotation See
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line 9 score
lines 9—7 annotation very important Puvis
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line 6 score
lines 6—4 annotation Read
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bottom-margin annotation vertically crossed in dark pencil To get titles it will be necessary to look over these notes at beginning of these notes, for it is impossible to make out by index th titles.
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lines 11—5 score
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line 5 score
line 5 annotation Sageret read
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line 2 score
line 2 annotation Herbert
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line 1 score
bottom-margin annotation (Read)
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line 3 score
lines 2—3 annotation Read?
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line 5 score
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lines 15—16 score
lines 14—17 annotation Theophrastus on crossing plants : how old!

line 24 score
lines 21—24 annotation I dare say V. Baer quoted by Lecoq in    V. Berg
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line 25 score
lines 25—26 annotation on Variation of Iris
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line 4 score
bottom-margin annotation Read Link on relation of grafting to Hybridisation

line 2bottom-margin annotation X----Puvis (Read)—?? ---- I have got impression that Puvis no good authority


line 20 annotation ✔ Read ?
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line 16 annotation ----X Read
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line 7 annotation in dark pencil Diel

line 18 score
lines 17—20 annotation Puvis on crossing of grasses
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line 1 score
line 2bottom-margin annotation X { Beitrage on pollen & Stigma being ready at different times


line 2 annotation Read
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line 18 score
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line 10 score


line 2 score in reddish-orange crayon


line 26 underline "Wahlverwandtschaft"
bottom-margin annotation ⧟ “Elective affinity” Chemical term Dict.