The two genetic species currently called Anopheles coustani species A and B are named Anopheles (Anopheles) coustani Laveran and Anopheles (Anopheles) crypticus n. sp., respectively. A neotype for An. Coustani is designated from chromosomally identified material from Madagascar, the original type locality. Polymorphic inversions and differences between An. crypticus and An. coustani are shown on a chromosomal map compiled from the neotype family.