156 ON THREE FORMS OF CATASETUM TRIDENTATUM. 
We thus see that every single detail of structure of the male 
pollen-masses, with some parts exaggerated and some parts slightly 
modified, is represented by these mere rudiments in the female 
plant. Such cases are familiar to every observer, but can never 
be examined without renewed interest. 
We now come to the third form, Myanthus barbatus, often 
borne on the same plant with the two preceding forms. Its flower, 
in external appearance, but not in essential structure, is the most 
different of all. It generally stands in a reversed position, com- 
pared with Catasetum and Monachanthus—that is, with the labellum 
downwards. The labellum is fringed, in an extraordinary manner, 
with long papille; it has a quite insignificant medial cavity, at 
the hinder margin of which a curious curved and flattened horn 
projects. The other petals and sepals are spotted and elongated, 
with the two lower sepals alone reflexed. The antenne are not so 
long as in the male C. tridentatum, and they project symmetrically 
on each side of the horn-like projection at the base of the labellum, 
with their tips (which are not roughened with papille as in the 
male flower) almost entering the medial cavity. The stigmatic 
chamber is of nearly intermediate size between that of the male 
and female forms ; it is lined. with utriculi, charged with brown 
matter. The straight and well-furrowed ovarium is nearly twice 
as long as in Monachanthus, but is not so thick where it joins 
the flower; the ovules are not so numerous as in the female form, 
but are opake and pulpy after having been kept in spirits, and 
resemble them in all respects. I believe, but dare not speak 
positively as in the case of the Monachanthus, that I saw the 
nucleus projecting from the testa. The pollinia are about a 
quarter of the size of those of the male Catasetum, but have a 
pertectly well developed disk and pedicel. The pollen-masses 
were lost in the specimens examined, by me; but fortunately M. 
Reiss has given, in the * Linnean Transactions,’ a drawing of them, 
showing that they are of due proportional size, and have the proper 
folded or cleft structure ; so that there can hardly be a doubt that 
they are functionally perfect. As we thus see that both the male 
and female organs are apparently perfect, Myanthus barbatus may 
be considered as the hermaphrodite form of the same species, of 
which the Catasetum is the male, and the Monachanthus the female. 
Tt is not a little singular that the hermaphrodite Myanthus 
should resemble in its whole structure much more closely the 
male forms of two distinct species (namely C. saccatwm and, more 
especially, C. callosum) than either its own male or female forms. 
