14 SIR DIETRICH BRANDIS—AN ENUMERATION 
Of the 6 ovules, as a rule one only develops into a perfect seed. 
At present three exceptions only are known. Two seeds in one 
fruit have been found in Dipterocarpus condorensis, Pierre, alatus, 
Roxb., and Dryobalanops aromatica, Gaertn. f£. The seed fills the 
entire fruit-cavity. As the one favoured seed grows, the dissepi- 
ments, which are often very thin, are pushed aside or torn, but 
the placenta remains attached to the base of the fruit-cavity, it 
becomes hard and woody, and with the remains of the dissepi- 
ments is, as a rule, enveloped by the testa and cotyledons. The 
5 abortive ovules increase a little, they become hard and shining, 
and in most species may be seen attached to the placenta at the 
apex. Sometimes, as in species of Dipterocarpus, the abortive 
ovules are on the outside of the seed in its upper part, attached 
in a circle to the testa. 
In its structure the seed shows great variety, and it may at 
once be stated that in this order the structure of the seed does 
not afford any certain generic characters. However, most species 
have one point in common—the cotyledons have long petioles 
when the seed germinates, and in many cases these petioles are 
well developed in the seed before germination. 
As may be concluded from the position of the anatropous 
ovule, the radicle is superior, that is its tip is directed towards 
the apex of the fruit. The radicle, or, more correctly speaking, 
the hypocotyl, varies in length and is often as long as the 
embryo. In this case the point where the cotyledons are 
attached to it is situated near the base of the fruit-cavity. The 
cotyledons are often unequal in size, one being larger than the 
other; in the majority of cases they are bifid to the base, that is 
to the point where they are attached to the hypocotyl. In many 
species they are divided into numerous lobes. Very often they 
are auriculate at base, and when the radicle is short it is included 
between the auricles. 
A considerable number of species have albumen in the mature 
seed. In these cases the embryo is small, the cotyledons are flat 
and thin, more or less folded, and sometimes lobed. Albuminous 
seeds are known from several species of Dipterocarpus, Hopea, 
Vatica, and Stemonoporus, and it seems probable that they will 
be found in all large genera. Species with albuminous and ex- 
albuminous seeds are often closely allied and very similar. 
Regarding the strueture of the exalbuminous seeds, the reader 
must be referred to the detailed description under each species 
