16 SIR DIETRICH BRANDIS—AN ENUMERATION 
into numerous lobes, there is an outer, slightly concave cotyle- 
don, and they may be classed under this type. 
IV. The cotyledons are fleshy, bifid to base, and the four lobes 
are prismatic, 3-sided, the two inner faces flat, the third 
outer face rounded; the hypocotyl, which generally is 
nearly as long as the embryo, is imbedded in a groove on 
the inner edge of the 4 cotyledonar lobes. This type is 
found in 4 species of Balanocarpus and in 6 species of 
Shorea, belonging to the sections Brachyptera, Pinanga, 
and Mutica, also in Hopea globosa, nov. sp. (Plate II. 
fig. 25, Balanocarpus coriaceus). 
Two remarkable facts connected with the structure of the seed 
must here briefly be noticed. In Balanocarpus the testa, which 
in this genus is thinly membranous, loosely enveloping the seed, 
seems to be adnate to the inner surface of the pericarp at the 
base of the fruit-cavity, at the foot of the persistent placenta. 
In Stemonoporus there is at the base of the fruit-cavity a 
remarkable fleshy or fibrous cup-shaped excrescence, Heim’s 
“cupule chalazique." Both facts must remain obscure until 
light can be thrown upon them by the study of fresh or alcohol 
material in different stages. 
The non-nitrogenous substances stored in the seed are starch 
and fat. So far as known, starch prevails in Dipterocarpus, 
Doona, and Vatica; fat in Pentaeme and Isoptera. In other 
genera it varies. Dryobalanops aromatica, starch; D. oblongi- 
folia, fat. Hopea odorata, starch; H. ferrea, fat. Shorea 
robusta and obtusa, starch; S. Gysbertsiana, Pinanga, stenoptera, 
aptera, hypochra, fat. 
Anatomical Characters. 
The peculiar anatomical characters of Dipterocarpacee have 
long ago attracted the attention of Botanists, and there is now a 
considerable literature on the subject, the more important publi- 
cations being the following :—Konrap Mtturr, “ Anatom. Verh. 
der Dipterocarpaceen,” in Engler’s Bot. Jahrb. ii. p. 446 (1882). 
— Pu. van Tizenem, “ Canaux sécréteurs des Plantes," Ann. Sc. 
Nat. Septième série, Bot. i. (1885), Diptérocarpées, p. 59.— 
SOLEREDER, Systemat. Wert der Holzstructur (1885), Diptero- 
carpeen, p. 81.—Bvmcx, “Sur les Diptérocarpées des Indes 
