NEW GENUS OF ORCHIDEA. 199 
invisible, except through the thin rostellum. On a back view of 
the column, the anther is seen to be firmly attached to the 
rostellum. It consists of two granular loculi quite separate from 
each other and dehiscing laterally (very early, for on opening a 
bud I found that they had already dehisced). They then form 
two cups, in which the pollinia lie transversely, andffall out on 
either side of the rostellum. The anther is attached to the 
column by a short but distinct filament. The pollinia are four 
in number, bright yellow, elliptic and curved, of a somewhat 
waxy, and certainly not granular, consistency. They have no 
appendage of any kind, and resemble most those of a Liparis or 
some such plant, and, as I have said, lie horizontally in the loculus, 
that is, with the long axis of each pollinium at right angles to the 
axis of the column. As they do not touch the rostellum at any 
point, it would naturally be expected that they would have no gland, 
which is commonly supposed to be derived from the rostellum ; 
and this is the case. They seem to fall out of the loculi extremely 
readily ; and from the normal position of the column it appears 
. that the plant would have to depend for its fertilization upon the 
chance adherence of the pollinia to the sides of the back of some 
insect which comes to devour the soft purple cushion in the 
centre of the lip. 
At first I imagined that the two loculi might represent two 
distinct stamens, and that the trian gular body I have termed 
rostellum might correspond to the shield of Cypripedium. But 
on examining flowers treated with potash and glycerine, I found 
that there was only one fibro-vascular bundle to the anther, which 
runs up the back of the column, terminating abruptly between the 
loculi. 
The front of the column is traversed by two bundles, one on 
either side, which terminate at the level of the stigma; but I 
could see none extending into the rostellum. Nor are there any 
traces of the lateral stamens ofthe outer whorl, either in the form 
of stelidia or of bundles. And there are also no bundles repre- 
senting those of the inner staminal whorl. These, however, are 
commonly absent in Neottiem, to which section this plant appa- 
rently must be referred. . 
For though the pollinia are in texture very different from that 
ofa Neottiacian plant, and bear more resemblance to those of 
the Epidendres, yet the complete adnation of the anther to the 
back of the rostellum is only parallelled by the arrangement of 
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