Tae 
OF CYTINUS FROM MADAGASCAR. , 467 
portion of the perianth-segments, on the lower portion of the 
style, and on the perianth-tube are numerous multicellular 
cylindrical or club-shaped processes. 
A transverse section of a perianth-segment shows it to be 
composed of homogeneous parenchymatous tissue, the cells being 
roundish or pentagonal; the ‚upper and lower epidermis are 
similar in structure, and stomata do not appear to be present. 
At the base of the perianth-limb at the point of juncture with 
the perianth-tube there are usually about five fibro-vascular 
bundles in each of the perianth-segments ; these, however, 
radiate like a fan from the base upwards, and branch dichoto- 
mously, and just beneath the apex of the perianth-segments the 
termination of some twenty bundles may be counted. A case of 
apparently abnormal growth of the perianth was met with in 
one of the female flowers examined, one of the perianth-segments 
becoming trifid a short distance above the base. 
3. Andr&cium.—The parasite is unisexual, the separate clusters 
consisting only of one sex. The staminate flower is globose and 
about jan inch in diameter. The anthers, twenty or more in 
number, are sessile on the column, and are arranged in a verti- 
cillate manner. The column, which is solid at the base, in the 
upper part is hollowed out in the middle, and has the anthers 
arranged on the outside contiguous to one another in a regular 
manner. The anthers are bilocular, and dehisce extrorsely and 
longitudinally, the two loculi being parallel. The pollen is some- 
What peculiar, being compound ; it is composed of four cells, and 
the compound grain has a diameter of ‘03 millim. The position 
of the four cells varies; sometimes they are arranged in the form 
ofa tetrahedron, or they may be in the form of a pyramid. 
If a transverse section be made of the ring of stamens, the 
column will be seen to be composed of homogeneous parenchyma, 
with a simple series of 8-10 vascular fascicles. In the wall of 
the anther the external layer of cells is thickened, and fibrous 
cells are absent. The composition of the bundles is of a some- 
what simple character, the xylem element being represented 
Principally by spiral vessels, and the perixylem is composed of 
short thin-walled cells. The cells composing the parenchyma of 
the column have brownish-coloured contents, and give a decided 
dark blue-black coloration with solution of sulphate of iron, thus 
ak the presence of tannic, and probably other astringent 
acids, 
