182 THE LATE DR. NEWELL ARBER: CRITICAL STUDIES 
LEPIDOSTROBUS MAJOR (Brongn.). (PI. 8. fig. 20 and PI. 9. fig. 21.) 
Sporophylls detached. 
1822, Filicites (Glossopteris) dubius, Brongn. Sur Class. Végét. Foss. p. 232 (footnote), 
pl. 13 (=2), fig. 4. 
1828. Lepidophyllum majus, Brongn. Prodr. Hist. Végét. Foss. p. 87. 
1835, Lepidophyllum trinerve, Lindl. & Hutt. Foss. Flora, vol. ii. pl. 152. 
1836, Lepidophyllum glossopteroides, Govpp., Syst. Filic. Koss, in Nov. Act. Acad. Cies. 
Leop.-Car. vol. xvii. Supp. p. 431, pl. 44. fig. 3. 
1854. Lepidophyllum majus, Geinitz, Flora Hainschen-Ebersdorf. p. 55, pl. 14. figs. 12-14. 
1855. Lepidophyllum majus, Geinitz, Verstein. Steinkohlentorm. Sachsen, p. 397, pl. 2. 
fig. 5. 
1862. Fruchtblatt von Lepidophloios laricinus, Goldenberg, Flora Saraepon, Foss, Heft 3, 
pp. 94 & 45, pl. 15, tig. 5. 
1870, Lepidophyllum majus, Schimp., Traité Pal. Végét.vol. ii. p. 72, pl. 61. lig. 87; 
pl. 64. fig. 9. 
1871. Lepidophyllum majus, Feistmantel, Abhandl. K. Böhm. Gesell Wissen. VI, 
Folge, Band v. p. 31, pl. 1. figs. 2-3. 
1845-75. Lepidophyllum majus, Feistmantel, Palæontogr. vol. xxiii. p. 41, pl. 42. figs. 2-4 
(? fig. 1). 
1877, ? Lepidophyllum binerve, Lebour, Illustr. of Fossil Plants, p. 103, pl. 52. 
1901. Lepidophyllum (? Lepidostrobus) majus, Kidston, in Trans. Nat. Hist. Soc. Glasgow, 
N.S. vol. vi. pt. 1, p. 63, fig. 9. 
1911. Lepidophyllum majus, Kidston, in Mém, Mus. Roy. Hist. Nat. Belgique, vol. iv. 
p. 154. 
1912. Lepidophyllum majus, Arber, in Phil. Trans. Roy. Soc. ser. B, vol. ccii. p. 251. 
Diagnosis.—Sporophylls very large, up to 12 cm. long, rarely less than 
10 em. in length, and almost 1:5 em. broad, lanceolate, acuminate, with a 
prominent median nerve in the lower part and a very marked. groove above. 
Pedicel wedge-shaped, about 2 em. long and 1 em. across at its widest part. 
Remarks.—No examples of these sporophylls attached to an axis are 
known from Britain. Dr. David White, however, in his ** Fossil Flora of the 
Lower Coal Measures of Missouri " (Monogr. 37, U.S. Geol. Survey, 1899), 
has figured (pl. 60) a cone Lepidostrobus missouriensis, D. White, with very 
similar sporophylls apparently attached to an axis. The sporophylls are 
more oval-lanceolate (2 cm. broad) than in L. major. While the two species 
may be distinct, the American specimen appears to leave little doubt that the 
organs known as L. major were sporophylls and not leaves. 
No. 2076. An almost perfect sporophyll of this species from the Bristol 
Coalfield is figured on Pl. 8. fig. 20, nat. size. It measures 11 em. in length, 
and 1:4 em. across at its widest part. In the centre of the lamina there are two 
parallel and rather broad ridges rather distant from one another and separated 
by a shallow groove. The basal angles of the lamina are slightly eared. 
No. 3830. On PI. 9. fig. 21 part of the pedicel of a sporophyll of this 
species is shown, enlarged three times to exhibit the reticular network which 
