252 MISS H. BANDULSKA ON THE CUTICULAR STRUCTURE OF 
direction, in marked contrast with the corresponding cells of A. eacelsa, 
A. Cunninghamii, and A. Gépperti. The cells between two stomata of a 
series are markedly wider and shorter than those between two rows of 
stomata, though elongated in the same direction parallel to the long axis 
of the leaf. The number of vertical rows of epidermal cells between two 
linear series of stomata is variable, being from four to fourteen. 
Stomatal Apparatus.—The stomata are situated on the floor of pits roofed 
in by four or five or more somewhat irregularly grouped subsidiary epidermal 
cells. These disturb the symmetrical arrangement of the parallel rows of 
epidermal cells between the stomatal lines, and cause the epidermal cells in 
their neighbourhood to curve round them, the appearance presented by a 
stomatal line being that of a row of large beads on a thin string—each bead 
being a group of subsidiary cells, and the string, the connecting epidermal 
cells of the same linear row. The poral opening is a narrow oblong with 
bluntly rounded ends, between which the granular guard cells can be seen. 
The stomatal axes are not oblique. The long axis of each is parallel to the 
long axis of the leaf. The subsidiary epidermal cells are not all alike. Two, 
those parallel to the long axis of the pore, are elongated vertically and short 
horizontally. The two which close in the pore horizontally are not much 
longer in one direction than the other. It is evident that neither external 
form nor cuticular structure suggest a very close relationship between 
A. Bidwillii and A. Güpperti. 
Cuticular Structure of Araucaria imbricata (Pl. 21. figs. 21, 22). 
Stomata occur on both surfaces of the lamina, and are, as in the other species 
of Araucaria, situated on the floor of pits. The epidermis is extremely thick- 
walled. Stomata occur in very regular linear series, each series being sepa- 
rated from the next by three to seven rows of thick-walled cells. Each series 
is uniseriate, and the stomata are very close, a line of eleven stomata having 
no interspersed epidermal cells, and then the series is interrupted by two epi- 
dermal cells separating the eleven from another five stomata, these succeeded 
vertically by two epidermal cells and then nine stomatal cells, and so on. 
Stomatal Apparatus.—The subsidiary cell group is larger in all directions 
than the corresponding group of cells in A. Gépperti, and larger in all 
directions than the ring of cells surrounding the pore in A. Bidwillä. It is 
very considerably larger than the equivalent cell group in 4. excelsa. The 
poral axes are not absolutely parallel to the length of the leaf, but are slightly 
oblique, though much less so than in A. excelsa and A. Gépperti. The oblong 
pore is very thick-walled, elongated, and narrow ; through it, in some cases, 
the slightly open guard cells can be seen. 
Thus, when we compare the external characters and the relative dia- 
meters of the axes of the poral ring, and of the epidermal cells of the 
