254 MISS H. BANDULSKA ON THE CUTICULAR STRUCTURE OF 
Cuticular Structure of Sequoia gigantea (Pl. 21. figs. 23-25).—This has 
been described by Rudolph Florin (1920). He does not give detailed 
measurements of the size of the various cells, but merely the ratio between 
their length and breadth in each case. He describes the stomata as occur- 
ring on both surfaces of the leaf, and says that the space between the rows 
of stomata varies just as does the distance between each row. He says 
further, that they are orientated lengthwise and that their long axis is also 
parallel to the direction of the vascular bundle, and points out that the 
openings do not lie in definite parallel closely connected rows any more than 
do the cells surrounding them, but between the rows of stomata, on the con- 
trary, the epidermal cells lie in fairly regular vertical rows, and have an 
elongated form. The ratio of breadth to length varies from 1 : 10 to 1:4. 
The opening between the epidermal cells which surround the guard cells is 
lly broad and 31, long. 
Detailed characters of the Cuticle (Pl. 21. figs. 23, 24, 25). 
I find there is a distinctly banded arrangement of the stomata, two bands 
occurring on either side of the wide midrib, these are somewhat irregular, 
and the number of serial rows making up a band varies from two to five - 
according to the distance from the apex. At the extreme apex a band 
consists of a single linear series of stomata, while at the base the banded 
arrangement is difficult to recognise owing to the widening out of all the 
series and the increase in the number of rows comprising the band. The 
stomatal axes do not preserve an exact parallelism, and though many are 
vertically orientated, a considerable number are oblique to horizontal in 
direction. This is comparable with the orientation of the stomatal axes in 
Araucarites Göpperti, but there is relatively greater axial obliquity in the 
stomata of Araucarites Gépperti. The middle of the leaf is occupied by a 
wider zone of vertically elongated narrow cells with rather oblique cross-walls. 
On either side of this zone and between the stomatal series the parenchyma 
is less regular, and the cells are shorter and wider, and though some are still 
vertically elongated, others have their long axis horizontal. The pore is an 
oblong slit surrounded by a ring of five, occasionally four or six, epidermal 
cells. Through the pore formed by this ring, the guard cells can be seen. 
Each stomatal series consists of a varying number of epidermal cells, two to 
five or many intervening between successive stomata, which are usually not 
in very parallel lines. One, two, or three rows of epidermal cells intervene 
between each series, the number increasing towards the base. 
The average length of the poral opening is ‘022 mm. 
The average width of the poral opening is ‘009 mm. 
Hence ratio of length to width is 2 : 1. 
The subsidiary epidermal cells have an average length of (0309 mm., and an 
average width of ‘019 mm. 
Hence ratio of length to breadth is 2 : 1. 
