256 MISS H. BANDULSKA ON THE CUTICULAR STRUCTURE OF 
surface, This is due partly to the smaller size of the individual cells of the 
ring, and partly to the smaller size of the pores. The epidermal cells are 
distinctly shorter than those of the lower epidermis. 
Average diameter of short axis=‘022 mm. 
Average diameter of long axis—:269 mm. 
Kräusel (1920, p. 353) considers that the cuticular structure of S. semper- 
virens is very much like that of Sequoia Langsdorfü, and like the figures 
which Reid gives of S. Coutisieæ, but he does not seem to have examined 
Reid’s material. 
SEQUOIA Tournatu, Brongn. Gardner has described this from the Myrica 
Bed of the Middle Bagshot of Bournemouth, and states that although there is 
no direct evidence to connect it with any particular tribe of Conifers, it more 
nearly resembles Sequoia sempervirens than any other species. It is also 
very similar to Sequoia Langsdorfü. 
The following account is based on specimen V. 15103 (Brit. Mus., N.H.), 
the external form of which is figured in Gardner’s ‘ Eocene Flora,’ plate 5, 
fig. 12. 
External Characters.—Average length 1*6 em.; average width 2:25 mm. 
The foliage on one and the same branch may be imbricated and decurrent and 
much like that of Sequoia gigantea, or distichous and decurrent, much longer 
and wider and externally extremely like the foliage of Sequoia sempervirens. 
This is interesting in view of the fact that Miss Eastwood (1895) finds two 
different types of foliage on trees of Sequoia sempervirens: “ All large trees 
of Sequoia sempervirens have the upper foliage quite different from the lower, 
with intermediate forms,” 
Cuticle (PI. 21. figs. 28, 29, 30).—Both upper and under surface bear 
stomata in vertically elongated serial rows ; the pores are close together, 
separated by only one or two epidermal cells, while each vertical row of 
stomata is similarly separated from the next by one or two epidermal cells. 
Epidermal cells and pores are constructed on a much smaller scale than those 
of any of the known Sequoias examined, nor is there the sharp distinction 
between ring cells surrounding the pore and the remaining epidermal cells 
that is so characteristic of the Sequoias. The pores are all orientated verti- 
cally, the opening is a narrow oblong, its long axis vertically directed. The 
midrib is about 18 cells wide, and some of its cells, which are much shorter 
than those of the midribs of known Sequoias, have slightly sinuate walls. 
Stomatal Pores.—The guard cells cannot be seen, but the pore shows as an 
opening bordered by four epidermal cells, two closing it in laterally, and 
these appear to be those of the adjacent epidermal cell rows, slightly dis- 
tended, while the two which close in the pore horizontally are also vertically 
elongated and form continuations of the epidermal cells in the same linear 
series. Very occasionally a short, almost isodiametric cell is cut off to close 
in the pore horizontally. 
