CUTICLES OF SOME RECENT AND FOSSIL FAGACE.E. 427 
On the Cuticles of some Recent and Fossil Fagaceæ. 
By Herexa Baxpvrska, ARCS. MSc. LS. 
(PLares 39, 40, and 6 Text-figures.) 
Read 6th March, 1924.) 
Introductory.—In a previous investigation (Bandulska, 1923) of the 
cuticles of certain dicotyledonous fossil leaves from the Bournemouth 
Eocene, attention was concentrated on their structure, and their relation- 
ships were not discussed. The first part of the present paper deals with 
evidence for the inclusion of one of the described leaves, i.e. Dicotylophyllum 
Stopes Band., in the genus Nothojagus Blume based on a study of the 
cuticular structure of recent Nothofagus leaves. The material used in 
the investigation of the recent cuticles was obtained from Kew and the 
British Museum, and was treated in the manner described in my former 
paper. The second half of the paper describes the cuticular structure of 
various recent species of Fagus and records a fossil Fagus with cuticle 
preserved, from the Eocene of Bournemouth. 
I. Tug Genus Norzuoraavs. 
Nothofagus is a subantarctic genus confined to southern South America, 
New Zealand, Tasmania, and eastern Australia. Of the species studied, 
N. Dombeyi, N. betuloides, N. procera, and N.antaretiea are South American ; 
N. Solunderi, N. fusca, N. Menziesü, and N. Blairii occur in New Zealand ; 
N. Moorei is Australian, while N. Cunningham is found in Australia and 
Tasmania. 
The leaves vary in average length from 1:15 em. in N. Cunninghami to 
lb 5em. in N. Moorei ; their width ranges from *7 em. (N. Dombeyi) to 4 em. 
(N. Moorei). The margin is usually serrate, erenate, dentate or irregularly 
lobed, except in N. Solanderi, N. Moorei, and N. Blairi, where it is entire. 
The number of secondary veins on each side of the midrib may be as few as 
two or three (V. betuloides) or as many as seventeen (N. procera), and these 
make angles with the midrib ranging from 31° (N. Solanderi) to 52? 
(N. procera). For further details of habit, external characters, and distri- 
bution of ‚Vothafuyus, see Elwes & Henry 1908, Cockayne 1921, and Hooker 
1840, 1844, and 1852. 
Detailed Account of the Cutieles of the different Species of Nothofagus. 
NorBoFAGUs DomBeyi Blume. Lower Epidermis (Pl. 89. figs. 1, 2). 
The stomata are very definitely grouped into areas by small venules about 
5 cells wide with somewhat elongated oblique-walled parenchymatous cells, 
On the venules, especially at the angles where they meet, are multicellular 
