438 MISS H. BANDULSKA ON THE CUTICLES OF 
slightly tapering. Midrib strongly marked with eight lateral veins on each 
side. Range of angle made by lateral veins with midrib 37°-52°, Average 
angle 43°. 
Cuticular Structure (Pl. 40. figs. 35, 36).—The cuticle is delicate and readily 
over-macerated as in the living species of Fagus. 
Lower Epidermis.—Stomata are confined to the lower surface. They vary 
in size. The guard cells are broad and flattened at the poles of their long 
axes. They have inner thickened cuticular rims and are horizontally cuti- 
nized at each end. Their line of junction is not strongly thickened, and is 
easily destroyed by over-maceration. Four epidermal cells usually surround 
them. The cells constituting the epidermal parenchyma have sinuate walls ; 
the venules are about two cells wide, their walls being also sinuate, but 
the venule cells are narrower than those of the epidermis. Hair-bases are 
scattered over venules and parenchyma, and consist of a thick-rimmed basal 
collar, somewhat raised upon a number (about 8) of radially-disposed cells. 
The average diameters of the guard cells are ‘027 mm. by ‘017 mm. 
Upper Epidermis (see text-fig. 6).—This consists of sinuate parenchyma 
crossed by venules. There are hair-bases on the venules and among the 
parenchyma cells. 
TEXT-FIG. 6. 
Upper surface of Fossil Fagus from Bournemouth, 
SUMMARY. 
An investigation of the leaf of Dicotylophyllum Stopese shows that it must 
be referred to the family Fagaceæ, and included in the genus Nothofagus as 
distinct from Fagus. Hence it is suggested that it be called Nothofagus 
Stopesæ (Bandulska). 
Both the external form and the cuticular structure of the fossil named Fagus 
bournensis indicate its close connection with Fagus rather than with Notho- 
fagus. The general external characters of the Bournemouth species are in 
striking agreement with those of Fagus sylvatica. The cuticle is more hairy, 
has somewhat smaller stomata, narrower venules and smaller epidermal cells, 
and is more xerophytic in habit. It is in all probability a specifically distinct 
form, and I therefore propose to call it Fagus bournensis. 
