446 MR. R. D'O. GOOD ON THE 
When the radicle is established the upper part of the hypocotyl, which 
has meanwhile been slowly elongating, undergoes another and reverse 
curvature and becomes vertical, bearing the endocarp on its tip. The 
cotyledons are still inserted in the fruit and absorb the thin layer of endo- 
sperm, which contains reserve food in the form of alemone and fatty oils 
(Fauth, loc. cit). After the straightening of the hypocotyl, the cotyledons 
begin to elongate towards their bases and to emerge from the fruit until only 
the tips are imprisoned. At length the fruit is completely levered off, 
apparently by the divergence or spreading of the cotyledons, and falls clear 
Via. 3. 
A-C. Later stages in germination of Hippuris. 
of the seedling. The two cotyledons now spread out at right angles to the 
axis of the young plant. From this point the vegetative axis of the mature 
plant develops in the axil of the cotyledons, and the first leaves appear. 
These resemble the ordinary axial leaves and are borne in whorls, but are 
much smaller. They bear scattered stomata and a few of the peltate hairs 
so characteristic of the plant (see Solereder, ‘Systematic Anatomy of the 
Dicotyledons, Engl. ed. vol. i. p. 337). 
Glück (quoted on p. 443) states that the first whorl may have 2, 3, or 4 leaves, 
and that the second whorl has only 2 leaves, while the immediately subsequent 
