FORMS OF FLOWERS IN VALERIANA DIOICA. 99 
On the inflorescence © I, most of the flowers were of the types A & B, 
the styles ranging from 3:2-2-2 mm. in length, but one flower of the type D 
was found, whose style measured only 1:1 mm. 
On the head C II, the only four flowers in a mature condition were all 
within the limits of the type A, while on the inflorescence С III, four 
flowers were of the type А, two between A & D, and one of the type В. 
Similar, though less extensive, variations having been noticed in other 
plants, an analysis of several inflorescences was made, in an attempt to 
discover whether any phyllotactic or symmetrical arrangement of the 
different types of flower might be found to exist. Тһе flowers in Valeriana 
dioica are borne in a series of opposite axillary eymes ; in some cases there 
seems to be some suggestion that the shortest-styled flowers are generally 
borne in the lowest pair of cymes. In the spike referred to above as D I, 
long-styled flowers of the type D were present in all the cymes except the 
lowest pair. Of the flowers borne in this pair of cymes, only one approached 
type В; in the four other flowers the style reached respectively 1:2, 1:2, 1:0, 
and 1:2 mm. in length. That these were not simply young stages is shown 
by the fact that the anthers had dropped in each flower. 
Again, in the inflorescence C I, one flower of the upper cymes had a short 
style (l:1 mm.), but the average length of the style in the remaining six 
flowers was 2:8 mm. In the two mature flowers on the lowest суше the 
length of the style was only 2:2 mm.; and young flowers, of about the same 
age taken from the upper and lowest cymes gave, respectively, lengths of 2:0 
and 1:6 mm. 
In another plant, the flowers of one inflorescence were mostly of the 
type А, but two flowers of type В were found on the lowest pair of сушез; 
while on another inflorescence of this plant, in which most of the flowers 
were of the type D, a few found on the uppermost cymes were type А. 
On the other hand, in the inflorescence D Ш, no sort of arrangement 
could be discovered. One суше of the uppermost pair bore five flowers 
(in three of which the anthers had either withered or dropped), all of 
type E or D, the styles measuring from 0°4-0°6 mm. The corresponding 
суше on the other side of the axis bore three flowers, one near type А, with a 
style 2:6 mm. long, and two of type D in which the longer style only 
measured 0:9 mm. 
One other type of variation may be mentioned. In one plant all the 
flowers examined in 1905 had an unusually long corolla-tube below the 
spur (fig. 28). In these flowers the style measured from 18-2 mm. in 
length, and yet scarcely reached the level at whieh the stamens were inserted 
(fig. 28 a). In later years this elongation of the tube was less marked and 
the plant bore several flowers of the type A. 
