374 MR. T. F, CHIPP: A REVISION OF 
А Revision of the Genus Codonopsis, Wall. By T. Е. С\нїрг. 
(Communicated by W. Borrixe Hemstey, F.R.S., Е...) 
(With 4 Text-figures.) 
(head 19th March, 1908.] 
OwiNa to the material recently collected by Mr. A. Henry and Mr. Е. H. 
Wilson, it has become desirable to attempt a short account and enumeration 
of the species of the genus Codonopsis of the Natural Order Campanulaccie. 
Under the generic name of Codonopsis I have included Codonopsis of Wallich, 
as enunciated in Roxburgh’s * Flora Indica,’ ed. Carey, vol. ii. p. 105, 
Glosocomia of D. Don, in his * Flora Nepalensis,” p. 158, and many species 
which though obviously allied to both these genera could not be included 
under either of them. 
An alternative would have been to have kept the original Codonopsis and 
Glosocomia distinet genera, but this would have entailed the making of several 
new genera for those species not coming under either, which have been 
brought to light by the above-mentioned collectors. 
А comparison of the original descriptions of Codonopsis and Glosocomia 
shows these differences :—Codonopsis has sublinear filaments, а three-lobed 
stigma, and a three-celled ovary ; Glosocomia has glabrous filaments dilated 
at the base, setose anthers, two-lobed stigmas, and a five-celled ovary. The 
characters of the stigma and ovary have no diagnostic value, as they have been 
found to vary from 3- to 5-merous in the same species ; but with regard to the 
filaments, which is the only other stated difference, not only do the two 
distinct forms as given above occur, but also every intermediate stage between 
them. On these considerations, therefore, it has been thought best to group 
all the species together, retaining for them Wallich’s original name of 
Codonopsis. 
As to the position of the genus in its extended form:—Following the divi- 
sions of Campanulacece as given in Bentham & Hooker’s ‘Genera Plantarum,’ 
vol. ii. p. 557, it belongs to the tribe Campanule with the regular corolla 
and free stamens, and under the subdivision of those with dehiscent capsules, 
as opposed to those with indehiscent berries. The genera nearest allied to it 
are Platycodon of De Candolle, the only species of which is an erect, robust 
plant, with sessile leaves and very large flowers, while Codonopsis is a scandent 
or climbing herb, or at most only suberect with weak stems and nearly 
always petioled leaves ; Leptocodon of J. D. Hooker & Thomson, which differs 
in having five epigvnous glands ; and Сатрапитеа of Blume, which has 
* 
baceate fruit. 
When Wallich described the genus in 1824 only three species were known, 
and one of them was so unlike the other two that he says he was of the opinion 
it ought to have been put in Polemoniacese. He mistook the short sterile 
foliaceous branches for compound leaves, resembling those of Polemonium. 
