GARDENIA THUNBERGIA AND ITS ALLIES. 419 
small flowers and ribbed fruits of G. Aqualla, and the almost entirely 
suppressed overground stems of G. subacaulis and G. Tinnee. The two last- 
named species are interesting instances of adaptation of arborescent types to 
the peculiar edaphic conditions of the savannas. 
With our present knowledge of Gardenia it would be idle to speculate on 
the phylogeny of the “ Thunbergia” group. They certainly represent 
a distinct branch of the greater phylum of Eu-Gardenia, from which 
they must have become differentiated a very long time ago. We know 
nothing about the dispersal of their seeds except what can be inferred from 
the structure of their fruits and seeds, and the casual observations of the 
collectors. It seems that the fruits of some of them, at any rate, are eaten 
by elephants and antelopes, and probably other animals, evidently for the 
suceulent pulp in which the seeds are embedded. The seeds would in this 
way get into the intestinal canals of those animals, and might finally be 
carried to great distances from the mother plant ; but no means of transport 
across ihe seas suggest themselves, and it is evident that the Eu-Gardenias 
must have reached their areas, in so fur as they are separated by water, ata 
time when those areas were still connected by land, that is in pre-Tertiary 
times. 
In the following enumeration of the species of the “ Thunbergia” group 
we have confined ourselves to describing only those which are new or were 
imperfectly known, the key which precedes the enumeration including 
sufficient characters for the determination of the others. In the key we 
have placed together two acaulescent species for purely practical reasons. 
They have no doubt descended independently from each other, Œ. subacaulis 
probably being more intimately connected with G. asperula, and G. Tinnew 
with G. lutea and G. ternifolia. (Отто STAPF.) 
Key to the Species. 
Subacaulis; flos solitarius cum foliis in rosulam dispositis. 
Folia scabrida; corollæ tubus 4-8 em. longus .......... ee nn 1. subacaulis. 
Folia levia ; corollæ tubus circiter 35 em. longus. ....... TET 2. Tinnee, 
Arbores vel frutices. 
Corolla 8-9-mera. 
Ovarium 9-merum. 
Calycis tubus ultra 5 mm. longus, lobis oblongis ...... .. 8. Jovis-tonantis. 
Calycis tubus infra 3 mm. longus, lobis subfiliformibus 
demum reflexis ..... leet nnn е...... .... 4, Folkensü. 
Ovarium 3-6-merum. 
Calycis lobi spatulati, infundibuliformes. ..... T 5. Thunbergia. 
Calycis lobi oblongi (non spatulati). 
Calycis lobi lateraliter complanati, equitantes et 
decurrentes „еее ‚2...4. 6. Saundersie. 
Calycis lobi dorsaliter complanati, non decurrentes .. 7. asperula. 
