ME. a. BENTHAM ON TEltNSTIKEillACE^. 59 



pared rather to the insertion consequent on the enlarged torus in 

 Nymphaacea, Calycanthea, and others, than to the true calyci- 

 florous insertion, where the staininiferous disk is connate with the 

 calyx-tube beyond where it has separated from the ovary. We 

 therefore have no hesitation in incorporating Anneslea and Visnea 

 in the tribe of Ternstrcemiece proper. 



The Ixionanthece are made to consist of three genera, Ixionan- 

 thes, Ochthocosmus, and Pentaphylax, supposed to differ from Ca- 

 melliacece chiefly in their definite stamens. But, except in this 

 one respect, Pentaphylax has less affinity with Ixionanthes than 

 vvitli any one genus of true Ternstroemiacece. It is indeed so closely 

 connected with Eurya, that it can only be removed from its 

 proximity invito, natura. The habit, inflorescence, floral envelopes, 

 insertion of the stamens, ovary, seeds, and embryo are the same ; 

 and even in the number of stamens, we have some species of 

 Eurya itself in which it is reduced to five. The only differences 

 arc in the shape of the anthers (which is yet nearer to that of 

 Ternstrcemiece than to that of the other tribes), and iu the fruit 

 (less fleshy and loculicidally dehiscent, as in Gordoniece). We 

 therefore place it at the end of the tribe of Ternstrcemiece proper, 

 forming a connecting link between them and the Gordoniece. 



Ixionanthes and Ochthocosmus had already been indicated by 

 Planchon as forming a distinct group ; but he has never published 

 his views as to their affinities. The insertion of the corolla and 

 stamens in Ixionanthes is so decidedly perigynous — the persistent 

 corolla, the stigma, the embryo, the inflorescence, and other cha- 

 racters are so different from what we observe in Temstrcemiacea, 

 that its removal from that Order is no longer doubtful. Its affi- 

 nities must probably be sought among the Calyciflorous orders 

 allied to Saxifragece, and I therefore reserve their discussion for a 

 future occasion. Ochthocosmus, including the African species 

 lately proposed by Klotzsch as a distinct genus under the name 

 of Phylloeosmus (Kl. in Schonl. Nachl. p. 232, t. 2), is much less 

 perigynous ; but on the whole its connexion with Ixionanthes is so 

 obvious, that its general affinities must be governed by those of 

 the latter genus. It must be remembered also that the degree of 

 perigynous expansion of the staminal disk is very variable in all 

 the groups connected with Saxifrages. 



The Pyrenaricce of Choisy are supposed to consist of two genera, 

 Pyrenaria and C«/p«H<7ria,' distinguished from Gordoniece only by 

 their so-called bony seeds. These seeds are, however, really not 

 harder than in Camellia itself, to which genus Seemann has not 



