SEEDLING STRUCTURE IN THE LEGUMINOS#. 11 
band of secondary wood, but аге quite easy to distinguish therefrom by their 
slightly different reaction to stains. Four bands of phloem are present, ex- 
tending between the protoxylems. Very many large tannin-sacs are developed 
throughout the hypocotyl in the phloem and in the perimedullary zone, and 
also in the epidermis and two subepidermal layers. 
Immediately above the collet the metaxylem begins to break up into smaller 
groups; about twenty of these are found throughout the greater part of the 
hypocotyl. They are scattered fairly evenly all round the stele, and but for 
the gaps in the bands of phloem and pericycle fibres it would be impossible 
to locate the original positions of the root protoxylems. Each eotyledon takes 
two bands of phloem and fibres, and a long strip of xylem tissue containing 
many scattered small endarch bundles. 
The detailed behaviour of the vascular system is very difficult to work out, 
owing to the great division of the xylem groups, and because of the confusion 
caused by the tremendous development of tannin tissue. It seems fairly 
clear, however, that all the root xylem passes to the cotyledons, and that the 
plumular bundles do not contribute to the primary structure of the root. 
TRACHYLOBIUM HoRNEMANNIANUM, Hayne. 
Hypocotyl swollen at the base, rapidly narrowing to root. Cotyledons 
fleshy, ovate, with deep dorsal median longitudinal furrows. 
The features of transition agree to a large extent with those of Tamarindus 
indica. The transition is low: the tetrarch root has a large pith throughout 
and an almost continuous ring of xylem ; triads are formed at the collet, and 
the hypocotyl is stem-like. It was possible here to determine definitely that 
the whole of the root xylem connected directly with the cotyledons. 
T'annin-sacs are present in the phloem, and subepidermal secretory 
cavities. 
SARACA CAULIFLORA, Baker. 
Hypocotyl with one-sided collar at its base ; tapering rapidly to a long 
tap-root. Cotyledons thick, ovate, asymmetrical. 
- The seedling is of unusually tough consistency, owing to enormous develop- 
ment of. fibres in the perimedullary zone and between the phloem and endo- 
dermis. The details of the transition are extremely difficult to make out, 
owing partly to the fibres and partly to the deeply staining tannin-saes. 
There is little doubt, however, that the transition takes place in just the same 
way as in Tamarindus indica and. Trachylobium Hornemannianum, 
The root is tetrarch and the transition markedly low. It may be noted 
that Van Tieghem and Douliot (1888, p. 173) record diarehy in Saraca (species 
not named). 
INTSIA AFRICANA, /houars. 
The hypocotyl narrows rapidly to a long stout tap-root. Cotyledons fleshy. 
