SEEDLING STRUCTURE IN THE LEGUMINOSAJE. 21 
groups, and the lateral xylems tend to break up into a number of smaller 
bundles ; the fibres now form a continuous ring round the vascular tissue. 
Each cotyledon receives four main groups of xylem, which, however, are 
somewhat broken in pieces. 
Throughout the hypocotyl the phloem remains in four continuous bands, 
alternating with the planes of the root protoxylems: each cotyledon receives 
two such bands. 
The plumular bundles, six in number, appear at the cotyledonary node : 
the first two leaves are opposite in the intercotyledonary plane, and each 
takes three of these bundles. The plumular protoxylems do not contribute 
to the root protoxylems, these latter being entirely cotyledonary in origin, 
just as in the epigeal species. 
CHSALPINIA SEPIARIA, Aob. (РІ. 8. figs. 53, 54.) 
A woody climber. Hypocotyl very short, tapering to a long thick primary 
root. First epicotyledonary internode long, narrowing upwards: first two 
leaves opposite. 
A species closely allied to C. Suppan, and showing great resemblances in 
its seedling morphology, but of quite distinct anatomical structure, which 
rather approximates to that of Pithecolobium Unguis-cati. 
The root contains four equal stout plates of xylem surrounding a pith. As 
we ascend towards the cotyledons they all broaden out internally and assume 
a tangential position. Two of them enter the cotyledons; the other two pass 
up into the epicotyl and enter the first two leaves; the two systems of bundles 
being distinct both in proto- and metaxylems. 
A noteworthy feature is exhibited by the plumular bundles. At the 
cotyledonary node these have not yet assumed the normal *stem.like " 
appearance with endarch xylem, but each consists of a double band of 
vessels developed tangentially on either side of a mesarch protoxylem 
fig. 53). This condition slowly ehanges as we ascend the epicotyl, and 
10 mm. above the cotyledons the transition to the stem-structure is 
complete. The other epicotyl bundles are endarch from their insertion on 
the cotyledon traces. This occurrence of a hypocotyl-like structure in an 
epicotyl can hardly be regarded as other than a result of the direct connec- 
tion with the root xylems. The transition from root- to stem-structure is here 
incomplete below the cotyledonary node. Similar though less well-marked 
phenomena occur in other cases where the plumular xylem is in direct con- 
tinuity with the root protoxylems. (See, e. g., certain Viciez.) 
One seedling showed the peculiarity that one of its cotyledonary root-poles 
divided into a triad very much below the others (fig. 54). This, however, 
was without any great effect on the transition. 
