bo 
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MR. R. H. COMPTON: AN INVESTIGATION OF THE 
GUILANDINA BONDUCELLA, Linn. (РІ. 8. figs. 51, 52.) 
Climbing shrub. Very large seedling. Collet at the base of the cotyledon 
stalks. First epicotyledonary internode elongated. 
The structure is of the same general type as that of Cesalpinia Sappan, with 
the exception that the changes begin at a lower level (about 10 mm. below 
the node), this being doubtless correlated with the greater size. The proto- 
xylems become separated from the metaxylems, and lie in groups external to 
them ; they undergo divisions independently of one another (fig. 51, 5 mm. 
below the node; fig. 52, 2 mm. below the node). All the root proto- and 
metaxylems are cotyledonary, each trace being composed of four bundles. 
Stout lignified fibres bound the phloem in the upper part of the hypo- 
cotyledonary axis ; and the epicotyl has a continuous ring of them within 
the endodermis. 
PAPILIONAT A. 
SOPHORE-E. 
SOPHORA TETRAPTERA, J, Mill. (Pl. 3. figs. 55-60.) 
A small or middle-sized tree. Cotyledons hemispherical, opposite ; plumule 
arising sometimes immediately between them, sometimes laterally, and 
developing early. First two leaves often nearly opposite. 
The root contains a curious-looking diarch strand of xylem, oval in trans- 
verse section and consisting of a mixture of vessels and thick-walled elongated. 
storage parenchyma. About 12 mm. below the cotyledons this strand breaks 
transversely into two similar wedge-shaped masses (fig. 55) ; these rapidly 
broaden, and each becomes converted into a continuous stout V-shaped band 
(fig. 56) 8 mm. below the node. These bands then break into triads, each 
consisting of a median crushed protoxylem and two lateral bands of meta-. 
xylem ; the phloem also dividing as the stele elongates in the cotyledonary 
plane (fig. 57, at 4 mm. below the node). Fig. 58, at 2 mm. below the 
cotyledons, shows each metaxylem dividing into two groups (the section 
from which the figure was taken being oblique, the upper group of bundles 
belongs to a higher level than the lower) : of these two halves of each meta- 
xylem, one enters a cotyledon ; the other remains in the axis and becomes a 
plumular bundle, as indicated by the dotted lines. The phloems divide in 
a corresponding manner. Fig. 59 shows the structure just above the node. 
Each cotyledon trace consists of a pair of collateral bundles with endarch 
xylem and a small median protoxylem. The four plumular bundles are 
shown branching ; the four branches towards the intercotyledonary plane 
fuse in pairs, and six bundles are produced in the main part of the internode 
(fig. 60). The structure of the epicotyl is, however, subject to some 
variation. | 
