SEEDLING STRUCTURE IN THE LEGUMINOSJE. 37 
Hosackia PURSHIANA, Benth. 
Erect annual. Material much etiolated, so that full dimensions cannot be 
given. 
The root was very short in the nine seedlings examined, and in all cases 
exhibited four 1-2-seriate xylem plates : triarchy has, however, been recorded 
by Van Tieghem and Douliot for /Josackia (sp. *), and this may be the case in 
longer roots of H. Purshiana. The transition seems to resemble that of 
Anthyllis tetraphylla in some respects, though the bulk of the tissues is less. 
The changes, moreover, do not being till two-thirds up the hypocotyl; they 
аге incomplete below the node, and in fact remain so far into the cotyledons, 
the lateral bundles not becoming truly collateral. 
Lorus TETRAGONOLOBUs, Linn. 
Prostrate or ascending herb. Hypocotyl tapering very gradually to a long 
primary root. Cotyledons rather thick, ovate, slightly asymmetrical, con- 
spicuously jointed to a stout petiole of 2 mm., strongly nyctitropic. 
The root is tetrarch, containing a solid stout xylem star alternating with 
four phloems, each with a strong group of fibres. The fibres throughout the 
seedling are rather conspicuous. This root-like structure persists without 
change for more than half-way up the hypocotyl. Here, however, there is a 
slight tendency for the four metaxylem bands to separate, and also for the 
protoxylems to become detached from their respective metaxylems (a 
phenomenon also observed elsewhere, e.g., in Guilandina Bonducella). As 
we enter the upper half of the hypocotyl there appears a pith which enlarges 
rapidly, and causes the lateral xylems to broaden out and finally to divide in 
half ; the polar xylem remaining in compact centripetal bands: thus, three- 
quarters of the way up the hypocotyl six bundles are present—viz., the two 
polar and the four half-lateral xylems. Just below the node the polar xylems 
divide into triads, and are joined by the half-laterals; the phloems re- 
maining in the same position throughout the hypocotyl, with the result 
that two cotyledon traces are organised, each composed of a median proto- 
xylem and a pair of collateral bundles, one on each side. The transition 
is therefore best described as ** intermediate-hi gh.^' 
The development of the fibres, which though not great is yet more 
pronounced than in most Lotes, seems to aid in the support of this tall 
seedling with its root-like hypocotyledonary structure. It is very unusual to 
find so high a transition in so thick a hypocotyl. 
Lorus REQUIENI, Mauri. 
Erect annual herb. Hypocotyl tapering slowly to the root. Cotyledons ovate. 
The root is tetrarch, and the whole structure of the hypocotyl very closely 
resembles that in Z. Tetragonolobus, save in details. Тһе transition takes 
place in the same region, and the cotyledon traces are similarly constituted, 
