52 MR. R. H. COMPTON: AN INVESTIGATION OF THE 
becomes broken into detached pieces at the collet. There are produced four 
groups of metaxylem which rapidly become endarch, one under each phloem ; 
the two polar protoxylems remain in situ. In the upper half of the mature 
hypocotyl the polar protoxylems disappear completely, and the structure 
becomes absolutely stem-like. Each cotyledon takes two of the collateral 
bundles. 
DERRIS scANDENS, Benth. 
A tall liane. The hypocotyl tapers to a long primary root. The cotyledons 
are ovate, asymmetrical, and foliaceous. 
The root contains a solid tetrarch xylem star and four phloems with 
included fibres. А pith appears 2 mm. below the collet, and at the collet it 
has dilated so as to break up the xylem ring into four tangential bands, each 
with central protoxylem (cf. Bauhinia, Pl. З. fig. 43, &c.). Intercalated 
bundles of metaxylem also may appear between adjoining bands of root 
xylem: these are similar to the © Zwischenstrünge" found so frequently in 
the Phaseoleze (Dodel, 1869). The polar xylems remain in continuous 
tangential bands, without showing the division into triads, in the lower half 
of the hypocotyl. The intercotyledonary xylems divide in halves about half- 
way up the hypocotyl, and three-quarters of the way up the polar xylems do 
likewise: the adjoining halves of polar and lateral xylems, however, do not 
fuse with one another, but pass into the cotyledons separately ; each cotyledon 
thus receiving four xylem bundles as well as interealary bundles. Meanwhile 
the phloem, originally in four distinct groups in the root, fuses into an 
almost continuous ring half-way up the hypocotyl; higher up it again 
separates, and at the node each cotyledon receives one band of phloem over 
the two polar half-xylems, and one band over each of the two half-laterals, 
besides a band over each “ Zwischenstrang” when present. The bundles of 
the cotyledon trace tend to come into contact with one another at the node, 
but do not actually fuse. 
VICIEZE. 
The seedling structure of the Viciew has received a good deal of attention 
in the past : in fact it appears that they have been regarded as representative 
of the Leguminose as a whole, rather to the exclusion of other tribes. 
Mlle. Goldsmith (1876, p. 1) gavea very complete account of the anatomy of 
Vicia sativa. Gérard (1881, p. 351) examined the structure of Lathyrus 
odoratus, L. latifolius, and Ervum Lens : these he found to agree substantially 
with Vicia sativa, while differing in certain respects, particularly in the 
presence of a pith at the cotyledonary node, obliterated higher up ; but he 
placed a different interpretation upon the occurrence of medullary xylem in 
the first few internodes from that given by Goldsmith. In 1910 Tourneux 
published a somewhat incomplete account of Vicia sativa, Frei Lens, Pisum 
