ANATOMY OF THE GENUS SALICORNIA. 323 
The occurrence of the spiral and spicular cells which are found in many 
species will be dealt with later. 
The vascular supply of each leaf is provided for by the passing out of a 
single bundle from the central stem cylinder. Immediately the leaf-sheath 
is reached the trace divides into three equal strands (ft. text-fig. 3, 3); 
the median one passes obliquely upwards, divides freely, and forms the 
numerous veins of the free leaf-tip ; the two lateral bundles pass obliquely 
outwards and curve downwards, branching repeatedly as they do so, and 
form the anastomosing network of bundles lying between the palisade 
layer and the aqueous tissue of the foliar sheath. 
The small ultimate strands die out before the base of the internode is 
reached, so that the leaf-trace vascular system of one internode has no 
immediate connection with that of any other (text-fig. 4, p. 324). 
In text-fig. 4, A-D shows one-half of the vascular tissue belonging to the 
two leaves whose apices are shown at /7 and /2, the remaining half of the 
foliar bundle system cannot be shown in the diagram. B-C gives the com- 
plete vascular tissue of the leaf whose apex appears at l3, that of the opposite 
leaf of the pair, 74, lies in the plane of the paper. The ramifications of the 
foliar bundle /4 show no anastomosing with those of /2, so that the network 
of bundles in the sheath is not “closed” completely, as is stated by 
Solereder *, but is, as Duval-Jouve f. remarks, an “open” system. Тһе 
individual strands of the leaf, both in its free and in its adnate region, are 
very small and show * reversed orientation" in a morphological sense, as 
has been pointed out by Fron f; Dangeard $ makes the same statement for 
the “cortical bundles" of Arthrocnemum fruticosum, Moq.| Although the 
orientation of the bundles is morphologically “reversed,” the histological 
relations are normal, the phloem being directed towards the palisade, while 
the xylem lies nearest to the ventral surface (РІ. 15. fig. 2). The morpho- 
logical inversion of the bundle is brought about by the * rotation " of the 
strand derived from the outgoing leaf-trace, during its passage through 
the foliar sheath towards the palisade layer. 
3. The Structure of the Flowering Shoot. 
The flowering spike is constructed on the same general plan as the 
vegetative shoot. It consists of a series of decussately arranged bracts, in 
* Solereder, H.: Systematic Anatomy of the Dicotyledons. Trans. by Boodle and Fritsch, 
1908. 
T Bull. Soc. Bot. Fr. xi. (1866) p. 140. 
і Fron, G.: in Ann. Sci, Nat., sér. 8, Bot. t. ix. (1899) p. 157. 
§ Bull. Soc. Linn. Normandie, sér. IV. ii. (1887) p. 188. 
|| The Arthrocnemum fruticosum of Moquin-Tandon includes Salicornia perennis, Mill 
(258. radicans, 8m.). 
