ANATOMY OF THE GENUS SALICORNIA. 331 
and S. pusilla, though the vegetative shoots are generally destitute of them ; 
in S. prostrata var. Smithiana they are found only in the reproductive spikes. 
Seedling plants of S. perennis, its variety lignosa, S. prostrata var. Smithiana, 
and S. ramosissima have usually no spiral cells, in the first. two species they 
do not usually appear to develop in the first year of growth at all *. 
Finally, in 8. Emerici f, Duval-Jouve [ = herbacea] 1, that author states 
that the spiral cells are few and ill-formed ; we are entirely in agreement 
with the first statement, for the elements are often absent in the vegetative 
shoot, yet when present we find that they show no difference in formation 
from those of other species. 
Spicular Cells or Stereides. 
In S. glauca, Del.$ spiral cells are absent, but they are replaced, as far as 
position is concerned, by stereides (Pl. 15. fig. 9). These are long, slightly 
branched elements with thick lignified walls, freely perforated by simple 
pits. They occur, as do the spiral cells, in the palisade region, and usually 
do not extend far into the aqueous tissue, they are extremely abundant, as 
are the spiral cells of S. fruticosa, Linn. 
According to Monteil ||, S. macrostachya, Moric. (= S. glauca, Del.) 
possesses both spiral cells and stereides in its palisade region, and he 
figures a specimen in which the latter occur mixed with short spiral cells. 
We have been unable to find this association of spiral and spicular cells 
in the same region, although we have examined material from four 
sources f. It is possible that Monteil worked on some form or species 
which was not available to us. 
In 5. glauca, Del. in addition to the stereides which occur so abundantly 
in the palisade region, there is an abundant development of similar elements 
round the vascular cylinder of the stem, just below the nodal region, and also 
in the free leaf-tips, parallel with, and just inside the epidermis of the ventral 
surface. Stereides are also abundant in the reproductive shoots. 
In S. fruticosa, Linn., the stereides are replaced by spiral cells in the 
palisade region, but stereides oceur in the other regions similar to those of 
5. glauca. 
In S. perennis, Mill., S. lignosa, Woods, and S. disarticulata, Moss, a few 
broad, short stereides occur in the reproductive shoots, especially at the base 
of the flower, while finally, in S. gracillima, Moss, the flowering spikes are 
abundantly supplied with stereides which are almost isodiametric. 
* In S. perennis only one spiral cell was found in all the first year material examined, 
T S: Emerici, Duval-Jouvez S. stricta, Dum. = S. herbacea, Linn. forma stricta, Moss, 
1 Bull. Soc. Bot. Fr. xv. (1868) p. 177. 
$ S. glauca, Пе]. = Arthrocnemum macrostachyum, Bunge. 
| Monteil, Anat. Comp. Feuille des Chenopodiacées, 1906, p. 130, fig. 41. 
4 Algeria, S. France, Palestine; British Museum Herbarium. 
