ANATOMY OF THE GENUS SALICORNIA. 333 
conclusion that they might be considered as * organes de soutien,” and stated 
that * elles remplissent alors le róle des cellules allongées à cloisons épaissies, 
qui constituent les cordons fibreux des Palmiers, des Pandances, ou qui for- 
ment le gaine externe du corps central de beaucoup de hampes florales." 
Van Tieghem * also considers that these elements are part of the cortical 
stereome. 
Holtermann T. describes the occurrence of spiral cells in Arthrocnemum 
indicum “die die Wasserzufuhr nach den peripherischen Teilen besorgen.” 
Volkens { and Solereder $ both state that they serve the purpose of 
water-storage. 
Finally Warming || expresses his opinion thus :—“ These water-storing 
tracheids seem to play the same róle as do wood-vessels in vascular bundles, 
since they fill with water and give it up again without collapsing.” 
Our own observations lead us to consider that the tracheids are water- 
storing elements, and their abundance in the foliar sheath at the nodal region, 
and in the reproductive shoots, is in accordance with this view, 
The funetion of the stereides needs no discussion, for they are obviously 
part of. the mechanical support of the plant. In their position and in their 
characteristics they closely resemble the similar elements found in the leaf of 
Tlakea suaveolens. 
Summary of the Distribution of the Spiral Cells and Stereides in 
the species of Salicornia. 
[. STEREIDES ONLY. 
S. glauca, Del. 
IT. SrEREIDES AND SPIRAL CELLS. (The latter only in the palisade 
region). 
a. Stereides in both vegetative and reproductive shoots. 
S. fruticosa, Linn. 
b. Stereides in reproductive shoots only. 
S. perennis, Mill. 
var, lignosa, Moss. 
S. disarticulata, Moss. 
S. gracillima, Moss. 
* Van Tieghem, P.: Traité de Botanique, p. 742. 
T Holtermann, C.: Der Einfluss des Klimas, 1907, p. 85. 
f Engler u. Prantl, Nat. Pflanzenfam. Teil iii. Abt. 1 a (1894), pp. 73-78. 
$ Syst. Anat. Dicot., Engl. Trans. 1908, ii. p. 1093. 
|| Warming: (1Zcology of Plents, 1909, p. 126. 
LINN. JOURN.—BOTANY, VOL. XLI. 2c 
