344 DR. E. DE FRA!'NE ON THE 
The tracheids and then the bundles end blindly in the aqueous tissue some 
distance below the apex of the leaf. Long stereides frequently occur below 
the epidermis on the ventral surface of the leaf, the long axis parallel with 
that of the leaf. | | 
Material for the investigation of the root and of the flowering shoot was 
unfortunately not available for examination. 
I should like to take this opportunity of thanking Professor F. W. Oliver, 
at whose suggestion this work was begun in the Field Laboratory at the 
Bouche d'Erquy, in Brittany, and in whose laboratory at University College, 
London, it was completed. 
A great part of the investigation was carried out at the University of 
London, Goldsmiths’ College, and many thanks are due to Mr. T. G. ШШ 
for his help and advice during its progress. 
We regret that the work of some investigators of the Salicorniew has had 
to be ignored. Some of the binomials which have been used are so vague 
that, until every single genus has been studied in detail and correlated with 
their work, it is impossible to ascertain which species they used in their 
investigation. 
SUMMARY. 
1. The succulent “cortex?” which covers the internodes of a Salicornian 
shoot is foliar in origin, and is phylogenetically derived from the basally 
developed leaf-sheath of the pair of leaves of the node above. This conclusion 
is based upon the evidence afforded by :— 
(a) The development of the shoot behind the apex. 
(^) The anatomy, and the course of the vascular bundles. 
(c) The structure of the flowering shoot. 
(d) The leaf-fall. 
(е) The morphology of the seedling. 
C/) The morphology of species of allied genera. 
2. The anatomy of the leaf is similar to that of any other succulent 
dicotyledon. It has a two-layered palisade parenchyma and well developed 
aqueous tissue. Numerous vascular bundles lie between the palisade layer 
and the water-storing tissue, The development of cuticle is slight, except 
in S. glauca and S. fruticosa. The structure of the free leaf-tips, the tubular 
leaf-sheath, and internodal sheath is practically identical. 
3. Each foliar trace divides into three on passing out from the stem at the 
node. The median strand branches, and forms the vascular system of the 
free leaf-tip, the two lateral strands curve outwards and downwards and 
by their repeated branching form the so-called “cortical” bundles of the 
internodal sheath. 
