DISTRIBUTION OF TRIBES. 419 
weeds in the Old World; the American species chiefly western 
and extratropical both in North and South America. ‘The three 
principal genera, Evax, Micropus, and Filago, are chiefly Old- World, 
but represented also by a few species in Western America; the 
sections Acantholena and Bombycilena of Micropus belong, how- 
ever, exclusively to the Old World; and the section Stylocline of 
Micropus, with the nearly allied genera Psilocarphus, Diaperia, 
and Micropus, are limited to California or Chili, or both. The 
somewhat more divergent genus /floga (Trichogyne, DC.) has a 
very different geographical area, being chiefly South-African, with 
one representative in the Mediterranean region. 
Three genera with many of the technical characters of Filagineze, 
but with a very different habit and geographical range, are not, 
perhaps, properly included in the subtribe, or at the most should 
only be considered its tropical considerably modified represen- 
tative. These are Cylindrocline, with one Mauritian species, and 
Blepharispermum and Athroisma, which together include four from 
tropical Asia or Africa. Their larger black achenes and reduced 
paleaceous or deficient pappus seem to bring them nearer to the 
Buphthalmex ; but the receptacular pales covering the filiform 
female florets are characteristic of Filaginee. They thus form an 
isolated group which would be still more out of place in any other 
tribe or subtribe with which they might be compared. 
Petalacte, a single South-African species, is another ambiguous 
genus which, technically, would be placed among Filaginee, but, 
from its habit, geographical, and some other characters, may be 
better considered as an exceptional Helichrysea next to Anaveton, 
which it so closely resembles. 
4, GNAPHALIEX, a very large subtribe, distinguished from 
Plucheineee chiefly by the scarious, or coloured, and radiating 
involucres, may be divided into two groups or large natural 
genera, Eugnaphaliee and Helichrysee, the one of about 200, the 
other of about 400 species, closely allied to and, as it were, passing 
into each other, insomuch that some botanists have united into 
one the two typical genera Gaaphalium and Helichrysum, but 
distinguished with few exceptions (chiefly tropical African) by a 
general character of some importance, the female florets outnum- 
bering the others and usually exceedingly numerous in the 
Eugnaphaliee, few, or disappearing altogether,in the Helichryses; 
and the geographical distribution is different. ^ Eugnaphalieze 
range over the whole world, forming only small distinct groups in 
