DISTRIBUTION OF TRIBES. 421 
with the involucre of a Gamocheta, but the pappus of a Eugna- 
phalium; and, 5, Anaphalioides, a remarkable New-Zealand group 
of two or three species, with the spreading involucres of Anaphalis, 
and in sexual arrangements connecting that genus with Gna- 
phaliwn. 
With regard to the groups so much further divergent from 
Gnaphalium as to have been retained as distinct genera, one, 
Achyrocline (fifteen species), is common to the New and the Old 
World; like many tropical-American (chiefly Brazilian) genera 
it has two or three representatives in tropical Africa. It is nearest, 
on the one hand, to the corymbose group of the section Eugna- 
phalium, but with narrow few-flowered capitula more densely 
corymbose, a Helichrysum-like involucre, &c.; and, on the other 
hand, to Stenocline-among Helichrysez, of which it has the aspect, 
but with the Gnaphalioid sexual proportions. Nearly allied to it 
is the monotypic tropical-African Chiliocephalum without any 
pappus. 
Chevreulia and Facelis are two small South-American Andine 
or extratropical genera connected with the Lucilia series of Gna- 
phaliwm of the same region, but differing, the one in the long beak 
into which the achene is produced, the other in the plumose 
pappus. Lasiopogon, from extratropical Africa south and north, 
and from the latter extending into the Levant, is another small 
genus, differing from some of the smaller Zugnaphaliee and 
Filagines of the same region in the plumose pappus. Phagnalon 
has a dozen species from the Mediterranean region more isolated 
in character. The anthers are often almost or quite tailless ; and 
the species were indeed formerly included in Conyza; but their 
involucres, habit, styles, and the anther-tails of some of the species 
justify their having been removed to Gnaphaliez. 
All the above genera have the disk-florets usually fertile as well 
as the females; in the following eight or nine genera, so nearly 
connected with Gaaphalium that they have most of them been 
united with it by some authors, the disk-florets are almost univer- 
sally sterile and often with undivided styles. Chionolena is 
Brazilian, allied in other respects to the American Gnaphalia with 
radiate involueres. —Luciliopsis is Andine, near the Andine Gna- 
phalia of the section Lucilia. Tafalla is also Andine, but more 
tropical, and in its habit and dicecious capitula connected with 
some species of Baccharis of the same country, but with the 
Gnaphalioid inyolucre, anthers, £c. Mniodes, again, another high- 
