422 MR. G. BENTHAM ON COMPOSITA,. 
Andine small genus with dicecious capitula, corresponds in other 
respects with Merope, also high-Andine. Antennaria and Leow 
topodon are mountain genera connected with the smaller Eu- 
gnaphalia, dispersed over Europe, Asia, North Ameriea, and the 
Andes of South America. Anaphalis, allied on the one hand to 
Antennaria, on the other to the corymbose Eugnaphalia, is inter- 
mediate between the two as to sexual characters, and differs from 
both in the usually radiating involucres. It is more Asiatic than 
any of the other genera, being represented in North America by 
only a single species out of about twenty-five. Stuartina, Demi- 
dium, and Amphidoxa, three monotypic genera from Australia, 
Madagascar, and South Africa respectively, are allied in habit to 
the small glomerate Eugnaphalia; but, besides the. frequent 
sterility of the disk-florets, they differ in the pappus reduced to a 
very few sete in Amphidoxa, entirely deficient in the other two. 
There remains only Raoulia, an Australasian mountain genus of 
fourteen species, chiefly from New Zealand, which may be said to 
be almost strictly intermediate between Eugnaphaliex and Heli- 
chryses, the proportion of the female and disk-florets being 
variable and often nearly equal, but certainly with a Eugnaphalioid 
tendency both in that and in habit. 
Helichrysez present one of those instances (such as Proteacee, 
Restiaces, &c.) in which a large very natural group of plants had 
spread over two regions, South Africa and Australia, now quite 
isolated, but then possibly in connexion with each other, in times 
sufficiently remote for them to have diverged in each region into 
different forms, and have multiplied greatly in both, without having 
preserved a single species in common. The Helichrysex, how- 
ever, have retained a closer affinity than the larger groups above 
mentioned. The South-African Ericacez have only a representa- 
tive order or suborder in Australia (Epacridee); Proteacew 
and Restiacee have tribes but no genera in common; among 
Helichryses there are common genera and even sections, but no 
species. 
Helichrysum itself, the largest genus of the subtribe, has, out of 
about 260 species, 137 South-African and about 60 Australasian 
(chiefly Australian with a few from New Zealand), and in each 
country has established distinct sectional races. The subgenus 
Lepicline is wholly South-African, as are also several sections of 
Euhelichrysum ; others of these are exclusively Australian ; but the 
sections Xerochlena and Ozothamnus, although chiefly Australian, 
