424 MR. G. BENTHAM ON COMPOSIT E. 
and two Brazilian. It is very nearly allied to Helichrysum itself 
but still nearer, in many respects, to the tropical African ash 
American Achyrocline, although the sexual relations place the two 
in different divisions of Gnaphalium. Gardner’s genus Leucopholis 
has two Brazilian species very closely allied to Stenocline, with the 
habit, however, of the American Chionolena, and differing from 
both chiefly in inflorescence, the few-flowered heads being closely 
sessile, and collected in a globular cluster or compound head, as in 
the Angianthez. 
The remaining genera of Helichrysex require but little notice 
to their geographical distribution ; they are all limited to South 
Africa or to Australia, without any of the forms sufficiently 
similar in their divergences in both countries to be considered 
representatives ; for the diverging characters are differently com- 
bined in the two regions. They are chiefly characterized by the 
involucres and pappus. South Africa has six of these genera, 
comprising sixteen species ; South Australia sixteen genera, com- 
prising forty-eight species. Among the latter I may particularly 
mention Millotia and Quinetia, both monotypic, as connecting in 
some measure the Helichrysex with the Senecionidez, having the 
peculiar almost uniseriate involucre so rare in Inuloidee, gener- 
ally so frequent in Senecionidem. Among the South-African 
genera the monotypic Phenocoma is remarkable for the foliage, 
which is that characteristic of the Relhaniee; but the filiform 
female florets and the broadly radiating involucres are rather 
those of Helichryse:e. 
5. The ANGIANTHEE proper constitute a group of eight genera 
and about sixty-four species, exclusively Australian, and, with the 
exception of two New-Zealand species of Craspedia, limited to 
Australia itself. With a Gnaphalioid habit and connexion, they 
are further removed from Gnaphalium itself than the Helichrysex, 
the capitula being always homogamous without any female florets 
whatever; the small closely aggregate capitula are also to be met 
with in some Filaginez ; but in the latter tribe there are always 
female florets embraced by the receptacular or involueral palez. 
And, geographically, the Filaginez are entirely wanting in Aus- 
tralia, the fatherland of the Angianthee. 
‘esulia, a monotypic East-Indian genus, stands alone. Its 
essential characters are indeed those of Angianthes; but its 
habit, its country, and several points of structure show but a re- 
mote affinity with any Australian genus of that subtribe. 
