DISTRIBUTION OF TRIBES. 429 
two South-African species; Bojeria one Mascarene and one 
South-African; and, lastly, Printzia, with five species, Homo- 
chete, with one species, and the two monotypic genera unknown 
to me, Alinurothamnus amd Cypselodontia, are South-African. 
Among all the above forms Porphyrostemma is the most excep- 
tional, having the purple, narrow, linear, very numerous ligulate 
corollas of Erigeron, with all the essential characters of Euinule:e. 
All the other genera are homochromous. 
There remains the somewhat anomalous genus Carpesium, with 
four or five species, which in its tubular female florets connects 
Euinulee with Plucheinee, but, upon the whole, is best placed in 
the former subtribe. Its geographical area is within the chief 
range of Euinulez, South Europe and temperate and tropical 
Asia. 
9. Bupnrustmes. The subtribe Buphthalmes, sixteen genera, 
but scarcely above fifty species, allied to Euinulee, has a nearly 
similar geographical distribution, somewhat more restricted east- 
ward, and offers some exceptions. Buphthalmex are chiefly 
African, European, and Oriental. Their structural connexions 
are more general than those of the other subtribes of Inuloidea, 
their styles the same as in Euinulee in the North-African and 
European genera, like those of the Athrixiez in some exclusively 
South-African forms ; and their tailed anthers and alternate leaves 
leave no doubt as to their place in this tribe; but in their rigidly 
paleaceous receptacle, the nature of the pappus in several genera, 
and some other respects they point to some connexion with 
Helianthoidex. The supposed affinity to Asteroidew appears 
more remote, and can only have been suggested by the numerous 
narrow yellow ligule of some genera. Amongst themselves, the 
genera, although small, are more distinct than most of those of 
the preceding subtribes. Of the seven most nearly connected 
with each other, three were long united under Buphthalmum; 
but two of the others have been hitherto placed in very different 
tribes, owing to inattention to the anther-tails, and also to the 
supposition that there were no Buphthalmes in South Africa, 
As it now stands, Buphthalmum is reduced to four species exclu- 
sively European: Odontospermum seven species, and Pallonis one 
species, formerly included in Buphthalmum, belong to the Medi- 
terranean region generally, the former extending to the Canarian 
and Cape-Verd Islands; but Callilepis, two South-African spe- 
cies, Sphacophyllum, one Mascarene species, and Anisopappus, two 
