496 MR. G. BENTHAM ON COMPOSITE. 
views. To gonofurther than the few figured by Hildebrand in 
his above-mentioned memoir in the ‘ Nov. Act. Nat. Cur.,’ what 
would he have done with the series Doronicum, t. 2. f. 23-26 
(an undoubted truncate penicillate Senecionida), Bidens, t. 1. 
f. 30, 31, Emilia, t. 1. f£. 11-13, Dahlia, t. 1. f. 26, 27, and 
Gaillardia, t. 3. f£. 1, 2, which last is as undoubted a Heli- 
anthaceous style according to his views, and passes into the 
Asteroid Solidago style, t. 2. f. 7-9? Where would he, where 
could he draw the line? And if he had gone through many of 
the common tropical genera, he would have been obliged to re- 
move Gynura far from Senecio, Dahlia from Coreopsis, Spilanthes 
from Verbesina, de. 
Delpino next divides Senecionide into two subfamilies, Sene- 
cionee, with zoidiophilous, and Artemisiacee, with anemophilous 
fertilization—according to him, a very remarkable and constant 
distinction, accompanied also by a difference in the position of the 
capitula, erect in one case, nodding in the other. As to the two 
modes of cross fertilization, or rather of the conveyance of 
pollen, it has hitherto been observed in so very small a number of 
species, that I must refrain from expressing any opinion as to 
their constancy as generic characters; but I would only refer to 
his own note, p. 34, as to the occurrence of the two modes in dif- 
ferent species of one and the same genus. Erect and nodding 
capitula occur not unfrequently (e. g. Lactuca and Prenanthes) 
in different species of the same genus; they are not constantly 
nodding in Artemisia; and the Melampodineous genus Parthe- 
nice, too closely allied to Parthenium to be widely separated from 
it, has the habit and nodding capitula of the Ambrosieous genus 
Cyclachena. 
Ambrosiex are remarkable for their anthers less perfectly con- 
nate than in any other Composite, although closely approximate, 
forming the usual cylinder and often slightly cohering ; they are 
also distinguished by their terminal appendages inflected or 
hooked at the end, as observed by A. Gray*. The anther- 
bearing florets are as constantly sterile as in Melampodinee and 
Milleriew, and the styles of these sterile florets as constantly 
* In a recent part of the ‘Proceedings of the Academy of Natural 
Sciences of Philadelphia, 1869, p. 189, Mr. T. Meehan observes that in 
Ambrosia artemisiefolia this inflexed setiform appendage is only to be found on 
anthers which do not present perfect pollen; the abundantly polliniferous 
anthers are broad, without horns. 
