a 
448 MR. G. BENTHAM ON COMPOSITA. 
of the American ones, is also found in Australia. If only a 
colonist there, it must be so ancient a one as to have undergone 
some slight modifications in form. Asa subtribe these Flaveriex 
are rather further removed from Helianthoides, and approach 
the Tagetinem ; their involucre is the most prevalent one m 
Senecionides, the style that of Anthemidez and the larger por- 
tion of Senecionidez, the achenes such as are prevalent in Bwries 
and Tagetinez. 
The S.-African monotypic genus Cadiscus is anomalous, but 
appears to me to be much more nearly connected with the Hele- 
nioid Flaveriex than with any S.- African type. 
The Tagetinee, 13 or 14 genera and above 100 species, form as 
a whole a very natural group, which will, moreover, very naturally 
divide into three, Porophyllum, Tagetes, and Pectis, taken each in 
the most extended sense. All three have their principal seat in 
the Mexiean region, but extend in a few species all over 
the warmer parts of South America; very few species reach Cali- 
fornia, none extend far eastward in N. America. A monotypic 
form diverging from Porophyllum (Lescaillea) is insular, limited 
to Cuba; another monotypic, Schizotrichia, is Peruvian. None 
are known from the Old-World except as introduced weeds, one or 
two species of Tagetes itself, long cultivated for ornament, having 
almost naturalized themselves in some parts of tropical Asia and 
Africa. 
As a whole the subtribe connects Helianthoide; with Sene- 
cionidez, Porophyllum and some species of Pectis having almost the 
pappus of the latter tribe. The whole, or nearly the whole, are re- 
markable for the large oily receptacles or glands scattered on their 
foliage and involucres. Pectis (40 species) has the style-branches 
much shorter than in any other genus of Helenioide:e, or of any of 
the nearly connected tribes, and is, moreover, marked by the rigid 
cilia at the base of the leaves or petioles. Syncephalanthus, a 
monotypic form included among those which diverge from Tagetes, 
has a very curious inflorescence ; the capitula are collected in 
clusters which assume precisely the aspect of the single radiate 
capitula of Bebera, the central capitulum of the cluster having 
no ray, and the ray-florets of the surrounding ones being only 
on the outer side, so as to form one continuous ray for the 
whole cluster. This peculiarity occurs also in the S.-African 
genus Cdera, and appears to have no special significance, syste- 
matical or geographical The genus Clappia, two species, so 
