DISTRIBUTION OF TRIBES. 461 
Arabian, and one Persian species, all nearly allied to the South- 
Afriean Othonna (from which they were first separated by Spach), 
but with the styles and involucres of Eusenecionez. 
Othonnez, excluding the two last-named genera, form a small 
subtribe of about 130 species in 5 genera, entirely S.-African, with 
the exception of Huryops, which has established one species in 
Abyssinia and another in Arabia. The three principal genera, 
-Euryops, 27 species, Gamolepis, 12 species, and Othonna (including 
Doria), about 80 species, appear to me to have a very close natural 
connexion, although usually placed in three widely distinct tribes. 
Euryops is generally acknowledged as a Senecionidea ; Gamolepis, 
only differing from it in the want of any pappus, has on that 
account been referred to Anthemidez, where it is evidently a 
perfect stranger; and an occasional absence of pappus in groups 
usually provided with it has now been observed in too many parts 
of the system to retain its formerly supposed importance. The 
third genus, Othonna, has been referred to Cynaroides, through a 
false appreciation of the style, which has neither the termina- 
tion nor the external ring of hairs or so-called articulation 
of that tribe, but is a genuine Senecionid style with a trun- 
cate penicillate tip; only, as the disk-florets are sterile, it 
remains undivided as in most other tribes under similar cir- 
cumstances. 
Liabezx is a small subtribe characterized by its imbricate invo- 
lucre and Vernonioid style. The principal genus, Liabum, of 
about 40 species, is S.-American, chiefly Andine, but extending 
in a few species into the W. Indies and northward to Mexico. 
It has been almost universally classed under Vernoniacese on 
account of its style; but the yellow heterogamous usually radiate 
capitula, as well as the habit, are very foreign to that tribe, whilst 
there is much that connects it with Senecionidew. The opposite 
leaves, though not common in the latter tribe, are to be met with 
in Arnica, Haploesthes, and Gynoxys; and the style is scarcely so 
far removed from that of Gynura as the latter from the ordinary 
truncate style of Senecio. The W.-Indian and Columbian genus 
Neurolena, two species, admitted on all sides to be a Senecionid, 
is very nearly allied to Liabum, and, indeed, closely connected 
with it through Schistocarpha, a Mexican and Peruyian genus of 
four species, with the opposite leaves of. Liabum and the paleace- 
ous receptacle of Veurolena. The small genera we have included 
in Liabum, differing from each other more in habit than in 
