DISTRIBUTION OF TRIBES. 463 
appears to me,on account of its yellow flowers and other characters 
to be referable rather to the present group, although it has even 
there no near connexions. 
The other extraneous genus is the S.-African Alciope, with two 
species, placed by De Candolle amongst Asteroidex, with which it 
seems to have much less connexion than with Senecionidezm. Its 
style is that of Cremanthodium ; the habit resembles that of Arnica, 
to which Thunberg referred it. It is not nearly allied to any 
genus of its own country. The anthers are those of the Sene- 
cionidez generally, and remarkable in that those of one species 
have the contiguous auricles of adjoining anthers connate, in the 
other the auricles are quite free and closed upon their own 
filaments. 
9. Calendulacee. 
Calendulaces constitute the smallest and most compact of the 
tribes we have adopted, and might almost have been enumerated 
amongst the subtribes of Senecionidez (with which it has much 
more affinity than with Cynaroidez, under which it is usually 
classed), but that there is a tendency to produce appendages or 
tails to the anther-auricles, and there is never any pappus. The 
sterility of the inner disk-florets, sometimes accompanied by a 
similar sterility in the ray, and the large size acquired by some 
or all the perfect achenes are also peculiarities, which justify the 
maintenance of the group as a distinct tribe. Itisalmost entirely 
African. Of the three largest genera, two (Dimorphotheca, twenty 
species, and Osteospermum, thirty-eight species) are exclusively 
S.-African ; Tripteris, twenty-eight species, is also S.-African, but 
has likewise a North-African subtropical or tropical species. Olt- 
gocarpus has three S.-African species and one in the island of St. 
Helena, whether aboriginal there or whether an introduction from 
S. Africa, and being yet to be discovered there, remains doubtful. 
Calendula, with scarcely ten species, although double that number 
have been described, belongs to the Mediterranean region, extend- 
ing from the Canary Islands to Persia. Dipterocome is a curious 
anomalous monotypic Persian genus, evidently allied to Oligo- 
carpus, but thus placed on the limits of the tribe both structurally 
and geographically. Eriachenium is another monotypic genus, 
which Schultz has correctly referred to this African tribe, although 
it comes from a distant land, Antarctic America. It is anomalous 
in babit, but nearer to Oligocarpus than to any other genus. 
LINN. JOURN.—BOTANY, VOL. XIII. 2L 
