The Geographical Distribution of the Frankeniaceae etc. 403 
3. Australia. 
Passing next to Australia we find an abundance and variety of forms 
sufficient to establish this continent as one of the three large centers of 
developement of Frankeniaceae, coordinate with the Mediterranean and 
South American regions. The species are allendemic and may be included 
in the genus Frankenia, although by no means do they fall within a single 
type, but on the contrary while developing a marked individuality, the 
Australian species fall into three groups, the first of which has its counter- 
part in the Mediterranean region and in Chili; the second in the isolated 
forms like F. triandra Remy of the Puna Region, and Beatsonia of St. 
Helena; and the third in Niederleinia of the salt-steppes of Argentine, and 
F. Palmeri and F. Jamesit of North America. 
Aspreviously stated, the grouping in general is that of »Pflanzen- 
familien« Subgen. Oceania stands appropriately for all of the species 
not found in the Mediterranean region. The genera Beatsonia and Nieder- 
leinia are to be regarded as to their origin as part of this subgenus. Then, 
instead of only the two sections Toichogonia and Basigonia, 1 would suggest 
a division of Toichogonia into Toichogonia cosmopolita to include all species 
of Oceania which correspond to section Eufrankenia of subgenus Afra, 
characterized by their much larger flowers and seeds; and Toichogonia- 
isolata to include all the remaining species of Oceania, which, although very 
widely differing in habit, are bound together by the one character of 
having very small flowers with ovules formed in the manner of Eufrankenia. 
As suggested in the name, the species in this section are isolated, and 
sharply defined, having evidently been separated from each other or from 
a common ancestral type for a very long time. 
The genus Beatsonia should be reckoned with this section. This leaves 
sect. Basigonia Ndz. unchanged but including also F. Drummondii Benth. 
and F. tetrapetala Lab. of Australia. The genus Niederleinia stands nearest 
this section. 
Toichogonia cosmopolita. 
F. pauciflora DC. occurs in numerous varieties in the characteristic 
habitats over the whole of the continent apd Tasmania‘), thus correspon- 
ding to the shrubby perennial type of Eufrankenia. But the most nearly 
allied species are F. nothria of Capland and the shrubby species of sect. 
Toichogonia cosmopolita in Chili. In F. pauciflora the flowers are larger than 
in any other member of the family except Hypericopsis, being even of greater 
length than those of H. persica, reaching 19—14 mm with calyx tube 
7—10 mm and 1!/;, —2 mm diameter. Seeds are produced in abundance as 
1) Benta., Fl. Austr. I p. 450. 
