CAMPANULACEOUS AJND OLEACEOUS ORDEBS. 9 



from Lobelia. Turning it over again and again, I can discover 



no plausible foundation for Delpino's genealogical Table from 



Lobelia to Artemisia. 



The present distribution of Campanulaceae seems to indicate a 



southern origin for Lobelieas and a northern one for Campanulese. 

 The southern extratropical or mountain-range of herbaceous 

 LobeliesB is extensive and varied. They are represented by iden- 

 tical genera, sections, or even species in South Africa and Aus- 

 tralia {Lobelia Bergiana, L. anceps, and allies), or in Antarctic 

 America, South-east Australia, and New Zealand (Pratia), and 

 have established several minor more or less endemic groups gra- 

 dually diverging from the common types in South Africa, Aus- 

 tralia, and Extratropical America. From thence Lobeliese appear 

 to have spread in several distinct directions into and beyond the 

 tropics, without any transverse northern connexion between the 

 several lines, which may be traced as follows : — 



First, and most abundantly, along the western mountain- 

 ranges of America, where they have developed into the shrubby 

 genera Siphocampylus, Centropogon, and Burmeisteria, of which 

 nearly two hundred species are already known, all remaining en- 

 demic in the tropical or subtropical mountain-regions, with the 

 single exception of the Centropogon surinamensis, which has gene- 

 rally spread over tropical America. Secondly into tall herba- 

 ceous true Lobelice with a wider general range, having formed, 

 however, special groups of a more local character, such as the 

 Tupce of Chili, the thapsoid Brazilian species, the Tylomia of the 

 West Indies, the Homochili of Mexico, the JEulobelice of extra- 

 tropical North America. Thirdly, into the Pratioid genera Hyp- 

 sela, Lysipoma, and BkizocepJialuni, which have remained almost 

 entirely confined within the South-American, Andine, or extra- 

 tropical regions, the JPratia hederacea alone, the exact counterpart 

 of the Himalayan P. begonicefolia, having extended further into 

 South Brazil. Fourthly, the southern Hemipogon group has main- 

 tained the typical characters in a few species thinly scattered over 

 the range, chiefly in Mexico, but has also developed into endemic 

 groups passing gradually into the Mexican Heterotopia, more ab- 

 ruptly into the north-western Downingia. The southern Lau- 

 rentia is also very closely represented by the Mexican L. ratno- 

 sissima and by the Californian Porterella, and the southern Iso- 

 toma by the West-Indian Hippobroma. 



In the Old World the northern dispersion of the Lobelieae ap- 



