88 MR. S. LE MARCHANT MOOEE ON THE OCCURRENCE 



a subglobular three-lobed and three-styled organ (fig. 17), the three 

 lobes of unequal size : the least bears a couple of small ovules ; the 

 next larger has two ovules of ordinary size (fig. 18), while in the 

 largest of the lobes are seated three stalked ovule-grotesques. 

 The knobbed appendages to the two longer of these stalks are very 

 far from being perfect ovules; but the point of greatest in- 

 terest relates to the small short-stalked body springing from the 

 sinus of dichotomy of the other two. ' This, as shown in the figure 

 (19), has a linear slit running the whole length across its anterior 

 face ; and appearances were so suspicious, that I carefully exa- 

 mined it for pollen ; and though I was not able to detect any, yet 

 I rather incline to the belief that it may be an incipient pollinife- 

 rous organ. 



I was able to examine only one more flower ; and this exhibits 

 a modification of form not before observed. In this, stamen No. 

 1 presents the appearance shown in fig. 16, but with the addition of 

 a third, lowermost ovule about half the size of the others. Stamen 

 No. 2 (fig. 20) is an unstalked pistilloid body with a short curved 

 hairy style, bearing inside its cavity, which communicates with 

 the outer world by means of a longitudinal slit continued to the 

 extremity of the style, three ovules, the uppermost of which is the 

 smallest. No. 3 has the same form as the last ; it contains four 

 well-formed ovules. No 4 is to all intents and purposes a carpel, 

 arising, as it does, but a slight distance below the organic apex, and 

 having a long curved style intertwined with the styles of the ova- 

 rian carpels. Its carpelloid nature is very emphatically pronounced, 

 inasmuch as it contains not less than five ovules, four of which 

 are full-formed, while the fifth is about half the size of the rest. 

 This organ is joined to the ovary about its centre by a narrow 

 isthmus of tissue, and has a completely impervious cavity. 



In all cases where an uncoiled style manifests itself, it is grooved 

 from base to apex, which condition also holds with the styles 

 of the two carpels composing the ovary of this last-mentioned 

 flower, one of whose carpels contains two, and the other three 

 ovules. 



I spoke above of a form having four hypogynous transformed 

 stamens and a three-lobed ovary ; and I venture to suggest that 

 this third ovarian lobe may be regarded as a reappearance of the 

 normally absent fifth stamen. This suggestion derives strength 

 from the fact that a modified stamen has undoubtedly been found 

 in organic connexion with the ovary ; and, on the other hand, if 



