STIGITATIC APPARATUS OF GOODEXOYIEiE. 205 



deep, and the stigma small and more buried in it. In Velleia the 

 mdusium is very large, broad, and, when closed, curved m a semi- 

 circle, but not deep ; the stigma does not appear to me ever to 

 protrude, but is never far from the orifice. In Calogyne Berardu 

 ana the style is split into two through the indusium and stigma, 

 each half of the indusium being a half-cup not joined on the 

 inner edge, but flattened and closely covering the half stigma, just 

 as if the Ooodenia indusium and style had been cut through with a 

 knife. In Calogyne piZosathe style is divided into three : instead 

 of being slit 3own the centre, the slit is on each side of the central 

 line ; the two lateral branches have each a portion of the indusium, 

 not joined on the inner edge, but closely pressed against a portion 

 of the stigma, as in C Berardiana ; but they have left behind them 

 the central portion of the indusium unjoined on both edges, but 

 closely embracing the centre of the stigma. That this division of 

 the style is not a separation of the carpellary leaves is proved by 

 this C. pilosa^ where the branches are three, whilst the carpellary 

 leaves (rudimentary cells of the ovary, valves of the capsule, &c.) 

 are, as in other Q^oodenovieae, two only. It would seem, there- 

 fore, as if it were a provision in some measure connected with the 

 duties of impregnation. 



In Bampiera the summit of the style, when short in the bud, has 

 the appearance of an ordinary peltate stigma, except that it is not 

 yet papillose, flat and nearly circular, wi^h the rudiment of the 

 stigma across the centre. It soon rises, the margins are raised into 

 a short almost 2-lipped indusium ; but I do not find that it carries 

 up any poUen with it ; and the stigma does not assume the perfect 

 appearance till the whole indusium and stigma has ensconced itself 

 between the two upper petals, which closely embrace it by means of 

 two thickened concave appendages, requiring some external agency 

 to open them and give access to the pollen. 



In Leichenmdtia we have another modification : the indusium is 

 usually described as broadly two-lipped, without any distinct 

 stigma. The fact appears to be that the upper less prominent Up 

 IS stigmatic all over, inside and out, with a transverse band of short 

 glandular hairs at its base outside, Avhilst the lower more prominent 

 lip is smooth and glabrous, or with a tuft of rigid hairs. Perhaps 

 this lower lip and the upper band of hairs are all that correspond 

 to the indusium of other genera ; and the so-called upper lip, out- 

 side of which impregnation may well take place, as observed by 

 Mr. Darv\nn, must be regarded as the true stigma. 



In giving these difierent modifications as characterizing the 



