84 MR. J. SCOTT ON THE FUNCTIONS AND STRUCTURE OF THE 
sules were produced, but these did not contain a single good seed. 
I may also state that, after inquiring of various cultivators of 
alpine plants, I cannot hear that this plant ever produces seed. 
In Primulas, Mr. Darwin has shown that the pollen of the “ short- 
styled” form, relatively to its own stigma, is considerably more 
sterile than the “long-styled” by its own-form pollen: may we 
not, then, in the “short-styled” forms of Gregoria Vitaliana have 
analogous relations of the sexual elements ? Anyhow, the above 
experiments render it highly probable that for the production of ` 
perfect fertility the conjunction of the two forms is absolutely 
necessary. 
4. The genus Cortusa is remarkable, though not peculiarly so, 
in apparently presenting the structure of the “long-styled”’ form 
only. I have at least failed in discovering the other form after a 
careful examination of the fine suite of specimens contained in 
the Edinburgh University Herbarium, as well as those in a few 
private herbaria. I find it to be the case also with cultivated 
plants of the C. Matthioli, the only representative of the genus ; 
and in the generic definition by Linnsus “ stylus filiformis, es- 
sertus” also occurs, so that it is highly probable that the one form 
only exists. I will now briefly describe the floral structure of the 
genus in so far as connected with the economy of fertilization. 
The corolla-tube is short, with the limb expanding upwards in 3 
campanulate manner ; the filaments short, scarcely half the length 
of the obeordately-acuminated, exeurrent anthers; the style is 
nearly three times the length of the stamens, and projects beyond 
the eorolla. A plant which I protected from insects, when in 
flower, did not produce a single seed: those, on the other hand, 
which I artifieially fertilized were perfectly fertile. I may fur- 
ther state, from the experience of others, as well as my own, that 
the plants of this species eultivated in our gardens, and left to the 
casual agencies of nature, produce in general a very small quantity 
of seed, —a consequence, perhaps, of their being less attractive to 
insects under cultivation than in their native haunts; for, un- 
doubtedly, perfect fertilization can only be effected by some such 
agencies. 
5. I have already stated that the structural relations of the sex- 
ual organs in Oortusa are not peculiar to it. In the allied genera 
Dodecatheon and Soldanella very similar relations are exhibited 
between the anthers aud stigmas, the latter projecting consider- 
ably, in the majority of the species, beyond the former,—an oecur- 
rence which, considering the systematic affinities of the three 
