104 MR. J. SCOTT ON THE FUNCTIONS AND STRUCTURE OF THE 
following comparative details of the results given in the table, 
In the first place we shall see, as to the relative degrees of fertility 
resulting from the homomorphie and heteromorphie unions in 
each of the crosses:—1. In the homomorphic unions of the long- 
styled Cowslip by pollen of the common Primrose the results are 
11 seeds per capsule, whereas from an equal number of flowers 
fertilized HETEROMORPHICALLY every seed-capsule aborted! 2. In 
the converse unions of these (the long-styled Primrose by pollen 
of the Cowslip) the results are singularly accordant, the homo- 
morphic unions yielding an average of 10 seeds per capsule, 
while in the HETEROMORPHIC wnions every seed-capsule is again 
abortive! 3. The heteromorphic unions of the long-styled Primrose 
by pollen of the Polyanthus yield an average of 26 seeds per cap- 
sule, and the homomorphie unions are utterly sterile. 4. Both 
unions of the short-styled Primrose by pollen of the Polyanthus 
are highly fertile; the heteromorphie also exceeding the homomor- 
phic unions in the proportion of 28 to 16 seeds per capsule—that 
is, as 5 to 3*. 5. And, lastly, we find that the heteromorphie 
unions of the short-styled Primrose by pollen of the Cowslip are 
remarkably fertile, yielding an average of 49 seeds per capsule; 
the homomorphie unions, on the other hand, do not yield a single 
seed ! 
We have already adduced a few illustrations of the functional 
relations of the two forms of one species with those of another 
species, and also expressed our belief, from the teachings of those 
illustrations, that the RELATIVE sexual powers of the two forms 
of a species did not extend to, or govern the results of wnions 
between the respective forms of two distinct species. How fully, 
then, are we supported in this view by the results given in 
the above table! how unequivocally do they show us that 
functional dimorphism is limited in its operations to the fn- 
dividuals of a species!t It is curious to observe how this is 
borne out by our experiments on the P. vulgaris and its modified 
descendants. Thus, in the above reciprocal unions of Primroses 
and Polyanihuses, the Aeeromorphic unions in both cases exceed 
* Mr. Darwin, in his letter to me accompanying the above results, remarks 
that the seeds of this short-styled Primrose were very small; so that we may 
perhaps suspect a number of them unfit for germination. 
`+ It is highly probable indeed, from the results of the red Primrose (vide 
Table V.), that we may yet have illustrations of the fertile unions of the modi- 
fied descendants of a species in which the laws of dimorphism may not be ob- 
served, and thus have cause to give i£ even a more restricted field of operation. _ 
