108 MR. J. SCOTT ON THE FUNOTIONS AND STRUCTURE OF THE 
short-styled forms, has also such functional characteristics as ren- 
der it a normal hermaphrodite representative of the species! 
We also see that the pollen-grains and stigmas of this non-dimor- 
phie form have become so differentiated with respect to their 
reciprocal action with those of the long- and short-styled forms, 
that a grade of sterility is induced greatly in excess of that which 
results from the homomorphic relatively to the heteromorphie 
unions of the latter forms! Now, Mr. Darwin has shown (and 
my experiments fully confirm it, vide Table IV.) that this lessened 
relative fertility of the homomorphic unions exceeds that resulting 
from the hybridism of several distinct species relatively to the 
pure union of those species. In view of such evidence, I think 
I am fully justified in adding that this non-dimorphie form is, in 
fact, judged by the physiological test so much insisted on by Prof. 
Huxley, a new and distinct species. Here we have an individual 
differing in colour, and likewise in important points of structure, 
from either of the forms which at present represent the species. 
Again, judged by the physiological test, we find a higher degree of 
sterility resulting from its unions with the two normal forms of 
the species, than that which results from the unions of other 
distinct species of Primulas! Certainly then, structurally and 
physiologically considered, this individual is perfectly entitled to 
specific honours. One class of naturalists would, indeed, readily 
admit the validity of such a claim if they had evidence of a con- 
stant recurrence of these characteristics ; but there is another class 
which consider ignorance of origin a necessary appanage to this 
claim ! 
The foregoing experiments and observations on the non-dimor- 
phic plant were made during its first or spring-flowering period, 
throughout which, as I have previously stated, a very constant 
equality was observed in the relative lengths of stamens and 
pistils in the flowers produced. A similar equality was observed 
in the relative lengths of these organs in the few flowers produced 
in its second or autumnal flowering; lately, however, it has 
produced an umbel in which the relative lengths of these organs 
in the respective flowers are so singularly variable that I have 
thought it worth while to give the measurements of each, as well 
as sketches of a few of the most distinct ; thus :— 
