THE THREE FORMS OF LYTHRUM SALICARIA, 173 
does not enclose any anthers. The end of the pistil is generally 
bent upwards at right angles. The six longer stamens, with their 
pink filaments and green pollen, resemble in size of the grains 
and in all respects the longer stamens of the mid-styled form, and 
both correspond in length with the long-styled pistil. The six 
shorter stamens, with their uncoloured filaments and yellow pollen, 
resemble in size of the grains and in all respects the longer 
stamens of the long-styled form, and both correspond in length 
with the mid-styled pistil. The capsules contain fewer seeds on 
an average than in either of the preceding forms, namely 83:5, and 
they are considerably smaller in size. In this latter respect, but 
not in number, there is a gradation parallel to that of the length 
of the pistil, the long-styled having the largest, the mid-styled the 
next in size, and the short-styled the smallest seed. 
From this description we see that there are three distinct 
female organs, or rather females as they are borne on distinct indi- 
viduals, differing in the length and curvature of the style, in the 
size of the stigma, and in the number and size of the seed. In the 
three forms, taken together, there are thirty-six stamens or males, 
and these can be divided into three sets of a dozen each, differing 
from each other in length, curvature, and colour of the filaments, 
in the size of the anthers, and especially in the colour and 
diameter of the pollen-grains. Each of the three forms bears 
half-a-dozen of one kind of stamens and half-a-dozen of another 
kind, but not all three kinds. The three kinds correspond in 
length with the three pistils: the correspondence is always be- 
tween half the stamens borne by two forms with the pistil of a 
third form. These remarks apply to the structure, and not, as yet, 
to the functions, of the reproductive organs. 
I ascertained the average number of seed by counting them in 
eight fine selected capsules taken from plants of the three forms 
growing wild, and the result was, as we have seen, for the long- 
styled (neglecting decimals) 93, mid-styled 132, and short-styled 
83. I should not have trusted this result, but I had a number 
of plants in my garden which, from their youth, did not yield the 
full complement of seed, but they were of exactly the same age 
and grew under exactly the same conditions, and were freely 
visited by bees. I took six fine capsules from each, and found 
the average to be for the long-styled 80, for the mid-styled 97, 
and for the short-styled 61. Lastly, I made numerous artificial 
unions, and, as may be seen in the following Tables, these gave 
in the long-styled an average of 90 seeds, in the mid-styled 117, 
