THE THREE FORMS OF LYTHRUM SALICARIA. 189 
the illegitimate unions are than those of the other two forms, we 
are led to consider the mid-styled form as eminently feminine in 
its nature. And although it is impossible to consider as rudi- 
mentary or aborted the two perfectly developed sets of stamens 
of the mid-styled form which produce an abundance of perfectly 
well-developed pollen, yet we can hardly avoid connecting, as 
balanced, the higher efficiency of the female organ with the lesser 
potency of the two mid-styled pollens. 
Finally, it is proved by the Tables that Lythrum salicaria 
habitually produces or consists of three females different in 
structure and widely different in function; that it produces or 
consists of three sets of males widely different in structure and 
function; and that two of the three sets of males are subdivided 
into subgroups of half a dozen each, differing in a marked manner 
in potency, so that regularly five kinds of pollen are elaborated 
by this one species of Lythrum. 
Lythrum Grafferi.—1 must now say a few words about some of 
the other species of the genus. I have examined numerous dried 
flowers of L. Grafferi, each from a separate plant, kindly sent 
me from Kew. This species, like L. salicaria, is trimorphic, and 
the three forms apparently occur in about equal numbers. In 
the long-styled form the pistil projects about one-third of the 
length of the calyx beyond its mouth, and is therefore shorter 
than in Z. salicaria; the globose and hirsute stigma is larger than 
that of the other two forms; the longer stamens, which are 
graduated in length, have their anthers standing just above and 
just beneath the mouth of the calyx; the half-dozen shorter 
stamens rise rather above the middle of the calyx. In the mid- 
styled form the stigma projects just above the mouth of the 
calyx, and stands almost on a level with the longer stamens of 
the previous form; its own longer stamens project well above the 
mouth of the calyx and stand a little above the level of the stigma 
of the long-styled form; the shorter stamens correspond in all 
respects with the shorter ones in the previous form. In the 
short-styled form the stigma of the pistil is nearly on a level with 
the anthers of the shorter stamens in the two preceding forms; 
and the longer stamens correspond with the longer stamens of 
the mid-styled form, and the shorter stamens with the longer 
stamens of the long-styled form. In short, there is a close general 
correspondence in structure between this species and L. salicaria, 
but with some differences in the proportional lengths of the parts. 
Nevertheless the fact of each of the three pistils having two sets 
