VASCULAR SYSTEMS OF FLORAL ORGANS. 177 
which is a lacuna formed by rupture. In the depression be- 
tween the buttresses a small portion of the style and conducting- 
tissue forms a sort of bridge. 
It is in this homogeneous mass of ground-tissue that we have 
a complete fusion of the hypertrophied borders of the carpels, 
which have thus entirely lost their individuality. The axis 
proper disappeared as soon as the tracheæ became disunited, as 
in figure 7 (i. e. it ceased to exist between figures 6 and 7). 
Hence the dotted lines in figure 9 radiating from the centre 
mark the ideal boundary of each carpel, and the line across the 
base of the ovary-cell shows the place where the rupture will 
take place when the fruit is mature. The column, or so-called 
“carpophore,” remaining, is therefore entirely carpellary in its 
origin *, 
xvii. Troræorum masus (Pl. XXVII.).—The pedicel of this 
plant, like that of Pelargonium, contains ten cords (1), which form 
a perfect circle. The cords are peculiar in having a zone of large 
cells between the tracheæ and phloém (La). Ten cords are given 
off to the calyx, the seven on the anterior side being united by 
transverse arcs (2). Five cords are given off to the petals in 
regular order (3, p.). Of the ten stamens, which theoretically 
belong to the flower, two, viz. one posterior and one anterior, 
are suppressed, the eight others receiving their proper cords (4). 
Three clusters of somewhat branching cords are isolated in the 
centre (4). Each of these first breaks up into five (5), all of 
each group being oriented towards a common centre, respectively. 
They increase to seven, surrounding the ovary-cells, with one 
much larger marginal cluster of tracheæ embedded in phloém (6). 
Each ovary-cell now becomes lobed round the cords. 
X. LEGUMINOS (Pl. XX VIL).—xrx. CYTISUS SCOPARIUS. 
The pedicel of the Broom contains a perfect and compact circle 
of cords (1). The first indication of an approaching irregularity 
consists of an unsymmetrical departure of three cords on one 
side (2) and then three on the other (3). Those on the posterior 
side become joined by loops. The central one branches, so that 
five cords are discovered (4). Three of these enter the circular 
base of the standard or vexillum (4, vex.). On the anterior side, 
besides the original three, two others are given off between them, 
* For further details, see ‘Origin of Floral Structures,’ p. 65. 
