VASCULAR SYSTEMS OF FLORAL ORGANS. 179 
As in others, the stamens are monadelphous at first (5), but 
diadelphous subsequently—the free stamen carrying two small 
wings, which have broken away from the attenuated edges of the 
tissue which binds the others together (6). 
In the formation of the pistil, it may be noted that, instead of 
there being three well-defined cords for the carpel, as occurs in 
so many plants, e. g. Hranthis (1v. 6-9), it is a complete circle 
which has to be utilized for one carpal (5). This would seem to 
show that there ought to be a whorl instead of only one carpei, 
as is of course always supposed to be the case, judging from the 
pentamerous pistil of Afonsea. It is interesting, therefore, to 
find that the distribution of the cords tends to corroborate this 
view ; four being arrested, their cords are utilized for one only. 
XXIII. CERATONIA Srrigua.—In the pedicel of the Carob, the 
cords are arranged in a somewhat quadrate manner (1). They 
then become circular, and send off five for the sepals and five 
for the petals (2). The ten for the stamens are given off in a 
circular form, when a perfect cylinder is left in the middle (3). 
This becomes oval for the pistil (4). When the ovary-cell 
appears, the two placental and the dorsal cords are of consider- 
able size, while a complete circle of cords surrounding the ovarian 
cavity is also developed. "These braneh and give rise to a cir- 
cumferential series (5). 
XI. ROSACEÆ* (Pl. XXVIL).—xxıv. Pyreus MaLus.—The 
Apple has a pedicel remarkably like a monoeotyledon, in that 
there are a number of cords indiscriminately placed ; but those 
on the circumference are the largest (1). In the case of the 
Daffodil, as in Anemone (it. 1), the larger cords are on the 
medullary side. These soon differentiate themselves into ten (2), 
five outer for the sepals (s.), and five inner for the petals (p.) ; anda 
star-like pentagon is formed out of the remainder, the lobes being 
superposed to the sepals (2, s.). The numerous stameus are formed 
by radial and tangential chorisis of the petaline and sepaline 
cords, as shown in figure 3. The lobes of the pistiline pentagon 
(2) break up into triplets (as seen in figure 4). One forms a 
* The remarkable feature of flowers of this Order is the presence of a well- 
defined and variously formed receptacular tube. P. van Tieghem has described 
and figured the anatomy of the tube of Prunus Lauro-cerasus (Recherches sur 
la Structure du Pistil, p. 40, pl. ix. figs. 20-24). I have fully discussed the 
nature of the receptacular tube, and the light thrown upon it both by Ana- 
tomy and Teratology elsewhere (‘Origin of Floral Structures, chap. x. p. 89). 
