VASCULAR SYSTEMS OF FLORAL ORGANS. 181 
into six placentary cords with central tracheæ (5), which take 
up their positions in pairs in front of the septa, and thence 
send off cords to the ovules (6); the tracheæ again being 
oriented as if axial. 
XIV. ONAGRACEÆ (Pl. XXVIIL).—xxvir. Fucusta.— 
In the example examined there were eight cords in the pedieel 
oriented asinfigurel. Alternate cords multiply until twelve are 
formed (2, 3, 4). The bases of the ovary-cells have not yet 
appeared. In the case described by Ph. van Tieghem, the ovary- 
cells appeared immediately after fig. 1. As soon as the ovules 
are reached, according to Ph. van Tieghem, each of the eight cir- 
cumferential cords are found to be doubled—the cords opposite 
the septa or sepaline giving rise to placental which send off 
branches along the septa, inwards; these then curve round to 
reach the base of the ovules. The dorsal and placentary cords 
are connecied by transverse branches. 
I did not, however, find this to be the case. It will be seen 
from tigure 2, that four cords (s.) are larger than the others, and 
that it is only the intermediate ones (p-) which multiply. The 
four which remain isolated are sepaline (s.); ana v isthe petaline 
which give rise to the innermost set of four placentary cords 
(3, pl.c.), one cord remaining as petaline (p.) ; these pass inwards 
and stand immediately in front of the sepaline (4). The ovary- 
cells now appear, and the petaline cords behind them give rise 
to the dorsal carpellary as well as staminal cords (5, st. d.c) ; 
but it is not until the summit of the ovary is reached that 
the sepaline and petaline give rise to the two whorls of stamens. 
At the top of the ovary the eight central cords become fused 
together, the four sepaline remaining free (6). This arrange- 
ment is then continued up into the style. 
XV. UMBELLIFER (Pl. XXVIII).—xx1x.  ANGELICA 
SYLVESTRIS.—ln this order the peduncle contains a definite 
number of cords (1) which form a circular network at the base of 
the umbel. This latier then throws off cords for each pedicel (2). 
At the base of the ovary five cords pass up the “ primary ” 
ridges of each mericarp from base to apex, while two more 
or less coalesce and pass up the middle. Figure 3 represents a 
section at the base of the ovary-cells, showing ten cords passing 
outwards to the circumference with two placentary cords in the 
