VASCULAR SYSTEMS OF FLORAL ORGANS. 183 
tube, which becomes somewhat pentangular, five are staminal 
(5, 6, st.). The other five increase by radial chorisis. The sec- 
tion of the tube now becomes more elliptical, and the anterior 
surface is clothed with papilla, showing that hypertrophy affects 
the anterior side much more than the posterior (7). 
It is noticeable that while the staminal cords have the tracheæ 
situated centrally (6, st.), the other or petaline cords are oriented 
as if they were axial. 
XXXIII. LEYCESTERIA FORMOSA (Pl. XXIX.).—The pedicel has 
eight or nine eords (1). These increase, giving off ten to the 
circumference, and retaining ten in the middle just below the 
appearance of the ovary-cells (2, ov.). As soon as these appear, 
five of the circumferential cords are situated dorsally, and five 
are intermediate (2). In the central space five of the cords stand 
opposite the septa (3); the others send off cords to the ovules 
and disappear (6, then 3). Higher up the axile placenta separates 
into five anchor-shaped structures, as seen in section, with a 
single cord in each each near the apex (4). At the summit 
of the ovary the central lacuna assumes a pentagonal form (5), 
aud ultimately closes up to form the style. The sepaline cords 
are remarkable for having arched girdles, from which are sent off 
cords to form the veins in each sepal (6). This reminds one 
of a similar process in Anemone (11. 2), in which the bracts of 
the involucre are supplied with their vascular system. See also 
the ale of Cytisus (xıx. 4, 6). 
XVIII. VALERIANE (Pl. XXIX.)—xxx1v. VALERIANA 
OFFICINALIS.—The pedicel has six cords symmetrically arranged 
(1); but they become unsymmetrical in the wall of the inferior 
ovary (2). On arriving at the summit two of them branch; 
these two branches, together with one of the original cords, enter 
the style (3), which terminates with three stigmas. Three of the 
other cords branch for the three stamens. The corolla has five 
of the original number. Fig. 4 shows a vertical distribution of 
the cords. 
CENTRANTHUS RUBER.—The arrangement of the cords is the 
same as in Valerian to the summit of the ovary; but a horizontal 
vascular ring now connects the whole of the cords. From this 
zone cords are given off for the corolla. It is observable that a 
corresponding ring, resembling the stipular zone in Galiwm, is 
LINN. JOURN.—BOTANY, VOL. XXVIII. P 
