VASCULAR SYSTEMS OF FLORAL ORGANS. 191 
XXVII. POLYGONACE (Pl. XXXI.).—r. RHEUM UNDU- 
LATUM.—The pedicel contains three or four principal cords (1). 
These increase and assume a triangular shape (2). Finally, it 
becomes six-angled in order to supply the six sepals. Three sepa- 
line cords give rise to three staminal; while six others arise in 
pairs, as shown in fig. 4. The remainder, three in number, formed 
from the three sepaline which kave not given rise to stamens, 
supply the carpellary cords (4). These finally result as dorsal 
carpellary cords, one having supplied a cord for the ovule at the 
base of the ovary-cell. 
XXVIII. ARISTOLOCHIACE (Pl. XXXI.).—11. Arıs- 
TOLOCHIA CLEMATITIS.—A section a: the base of the flower 
shows six cords passing off for the perianth or calyx, leaving six 
behind with a small lacuna in the centre (1). These six cords 
run up to the top of the ovary, then pass up the gynandrous 
column and bifureate at the level of the insertion of the anther 
(2, anth.), one branch passing off to each, the others continue up 
the style to the base of the stigmas (2, stig.). 
It has been thought that the styles and stigmas of the carpels 
are abortive in this genus; and that the now thickened con- 
neetive of the anthers, coherent laterally into a tube and covered 
above with stigmatic papillæ, play the part of styles and stigmas *. 
This idea is not borne out by a study of the anatomy, which 
seems to show clearly that it is only a case where a cord does not 
become differentiated into a staminal and dorsal carpellary until 
the level of the anthers is reached. The stigmas are therefore 
truly carpellary, as is apparently always the case in other flowers. 
Moreover, they are free from the filaments to a much greater 
degree in Asarum. 
XXIX. EUPHORBIACEE (Pl. XXXI).—ııu. EUPHORBIA 
Hetroscorra.—This has about ten cords in the peduncle below a 
three-flowered cyme (1). This at first forms a complete ring, and 
then throws off six cords, three on opposite sides (2); while 
small circles appear between the central one and the two groups 
of threes destined for the bracts (3). Thus three pedicels are 
formed. In the case where there are four flowers, one central 
and three circumferential, three cords travel outwards at each 
* See ‘Origin of Floral Structures,’ p. 83, note. 
